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Silurian

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Silurian
443.8 ± 1.5 – 419.2 ± 3.2 Ma
Silurian plate tectonics.png
Plate tectonics of Earth during the early Silurian
Chronology
Etymology
Name formalityFormal
Synonym(s)Gotlandian
Usage information
Celestial bodyEarth
Regional usageGlobal (ICS)
Time scale(s) usedICS Time Scale
Definition
Chronological unitPeriod
Stratigraphic unitSystem
First proposed byRoderick Murchison, 1835
Time span formalityFormal
Lower boundary definitionFAD of the Graptolite Akidograptus ascensus
Lower boundary GSSPDob's Linn, Moffat, UK
55°26′24″N 3°16′12″W / 55.4400°N 3.2700°W / 55.4400; -3.2700
Lower GSSP ratified1984[4][5]
Upper boundary definitionFAD of the Graptolite Monograptus uniformis
Upper boundary GSSPKlonk, Czech Republic
49°51′18″N 13°47′31″E / 49.8550°N 13.7920°E / 49.8550; 13.7920
Upper GSSP ratified1972[6]
Atmospheric and climatic data
Sea level above present dayAround 180 m, with short-term negative excursions[7]

The Silurian (/sɪˈljʊəriən, s-/ sih-LYOOR-ee-ən, sy-)[8][9][10] is a geologic period and system spanning 24.6 million years from the end of the Ordovician Period, at 443.8 million years ago (Mya), to the beginning of the Devonian Period, 419.2 Mya.[11] The Silurian is the shortest period of the Paleozoic Era. As with other geologic periods, the rock beds that define the period's start and end are well identified, but the exact dates are uncertain by a few million years. The base of the Silurian is set at a series of major Ordovician–Silurian extinction events when up to 60% of marine genera were wiped out.

One important event in this period was the initial establishment of terrestrial life in what is known as the Silurian-Devonian Terrestrial Revolution: vascular plants emerged from more primitive land plants,[12][13] dikaryan fungi started expanding and diversifying along with glomeromycotan fungi,[14] and three groups of arthropods (myriapods, arachnids and hexapods) became fully terrestrialized.[15]

A significant evolutionary milestone during the Silurian was the diversification of jawed fish and bony fish.[16]

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Ordovician

Ordovician

The Ordovician is a geologic period and system, the second of six periods of the Paleozoic Era. The Ordovician spans 41.6 million years from the end of the Cambrian Period 485.4 million years ago (Mya) to the start of the Silurian Period 443.8 Mya.

Devonian

Devonian

The Devonian is a geologic period and system of the Paleozoic era, spanning 60.3 million years from the end of the Silurian, 419.2 million years ago (Mya), to the beginning of the Carboniferous, 358.9 Mya. It is named after Devon, England, where rocks from this period were first studied.

Paleozoic

Paleozoic

The Paleozoic Era is the earliest of three geologic eras of the Phanerozoic Eon. The name Paleozoic was coined by the British geologist Adam Sedgwick in 1838 by combining the Greek words palaiós and zōḗ, "life", meaning "ancient life").

Rock (geology)

Rock (geology)

In geology, rock is any naturally occurring solid mass or aggregate of minerals or mineraloid matter. It is categorized by the minerals included, its chemical composition, and the way in which it is formed. Rocks form the Earth's outer solid layer, the crust, and most of its interior, except for the liquid outer core and pockets of magma in the asthenosphere. The study of rocks involves multiple subdisciplines of geology, including petrology and mineralogy. It may be limited to rocks found on Earth, or it may include planetary geology that studies the rocks of other celestial objects.

Silurian-Devonian Terrestrial Revolution

Silurian-Devonian Terrestrial Revolution

The Silurian-Devonian Terrestrial Revolution, also known as the Devonian Plant Explosion (DePE) and the Devonian explosion, was a period of rapid plant and fungal diversification that occurred 428 to 359 million years ago during the Silurian and Devonian, with the most critical phase occurring during the Late Silurian and Early Devonian. This diversification of terrestrial plant life had vast impacts on the biotic composition of earth's soil, its atmosphere, its oceans, and for all plant and animal life that would follow it. Through fierce competition for light and available space on land, phenotypic diversity of plants increased greatly, comparable in scale and effect to the explosion in diversity of animal life during the Cambrian explosion, especially in vertical plant growth, which allowed for photoautotrophic canopies to develop, and forever altering plant evolutionary floras that followed. As plants evolved and radiated, so too did arthropods, which formed symbiotic relationships with them. This Silurian and Devonian flora was significantly different in appearance, reproduction, and anatomy to most modern flora. Much of this flora had died out in extinction events including the Kellwasser Event, the Hangenberg Event, the Carboniferous Rainforest Collapse, and the End-Permian Extinction.

Dikarya

Dikarya

Dikarya is a subkingdom of Fungi that includes the divisions Ascomycota and Basidiomycota, both of which in general produce dikaryons, may be filamentous or unicellular, but are always without flagella. The Dikarya are most of the so-called "higher fungi", but also include many anamorphic species that would have been classified as molds in historical literature. Phylogenetically the two divisions regularly group together. In a 1998 publication, Thomas Cavalier-Smith referred to this group as the Neomycota.

Glomeromycota

Glomeromycota

Glomeromycota are one of eight currently recognized divisions within the kingdom Fungi, with approximately 230 described species. Members of the Glomeromycota form arbuscular mycorrhizas (AMs) with the thalli of bryophytes and the roots of vascular land plants. Not all species have been shown to form AMs, and one, Geosiphon pyriformis, is known not to do so. Instead, it forms an endocytobiotic association with Nostoc cyanobacteria. The majority of evidence shows that the Glomeromycota are dependent on land plants for carbon and energy, but there is recent circumstantial evidence that some species may be able to lead an independent existence. The arbuscular mycorrhizal species are terrestrial and widely distributed in soils worldwide where they form symbioses with the roots of the majority of plant species (>80%). They can also be found in wetlands, including salt-marshes, and associated with epiphytic plants.

Hexapoda

Hexapoda

The subphylum Hexapoda comprises most species of arthropods and includes the insects as well as three much smaller groups of wingless arthropods: Collembola, Protura, and Diplura. The Collembola are very abundant in terrestrial environments. Hexapods are named for their most distinctive feature: a consolidated thorax with three pairs of legs. Most other arthropods have more than three pairs of legs. Most recent studies have recovered Hexapoda as a subgroup of Crustacea.

Gnathostomata

Gnathostomata

Gnathostomata are the jawed vertebrates. Gnathostome diversity comprises roughly 60,000 species, which accounts for 99% of all living vertebrates, including humans. In addition to opposing jaws, living gnathostomes have true teeth, paired appendages, the elastomeric protein of elastin, and a horizontal semicircular canal of the inner ear, along with physiological and cellular anatomical characters such as the myelin sheaths of neurons, and an adaptive immune system that has the discrete lymphoid organs of spleen and thymus, and uses V(D)J recombination to create antigen recognition sites, rather than using genetic recombination in the variable lymphocyte receptor gene.

History of study

The Silurian system was first identified by British geologist Roderick Murchison, who was examining fossil-bearing sedimentary rock strata in south Wales in the early 1830s. He named the sequences for a Celtic tribe of Wales, the Silures, inspired by his friend Adam Sedgwick, who had named the period of his study the Cambrian, from the Latin name for Wales.[17] This naming does not indicate any correlation between the occurrence of the Silurian rocks and the land inhabited by the Silures (cf. Geologic map of Wales, Map of pre-Roman tribes of Wales). In 1835 the two men presented a joint paper, under the title On the Silurian and Cambrian Systems, Exhibiting the Order in which the Older Sedimentary Strata Succeed each other in England and Wales, which was the germ of the modern geological time scale.[18] As it was first identified, the "Silurian" series when traced farther afield quickly came to overlap Sedgwick's "Cambrian" sequence, however, provoking furious disagreements that ended the friendship.

Charles Lapworth resolved the conflict by defining a new Ordovician system including the contested beds.[19] An alternative name for the Silurian was "Gotlandian" after the strata of the Baltic island of Gotland.[20]

The French geologist Joachim Barrande, building on Murchison's work, used the term Silurian in a more comprehensive sense than was justified by subsequent knowledge. He divided the Silurian rocks of Bohemia into eight stages.[21] His interpretation was questioned in 1854 by Edward Forbes,[22] and the later stages of Barrande; F, G and H have since been shown to be Devonian. Despite these modifications in the original groupings of the strata, it is recognized that Barrande established Bohemia as a classic ground for the study of the earliest Silurian fossils.

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Origin of water on Earth

Origin of water on Earth

The origin of water on Earth is the subject of a body of research in the fields of planetary science, astronomy, and astrobiology. Earth is unique among the rocky planets in the Solar System in having oceans of liquid water on its surface. Liquid water, which is necessary for life as we know it, continues to exist on the surface of Earth because the planet is at a distance, known as the habitable zone, far enough from the Sun that it does not lose its water, but not so far that low temperatures cause all water on the planet to freeze.

Evolution of photosynthesis

Evolution of photosynthesis

The evolution of photosynthesis refers to the origin and subsequent evolution of photosynthesis, the process by which light energy is used to assemble sugars from carbon dioxide and a hydrogen and electron source such as water. The process of photosynthesis was discovered by Jan Ingenhousz, a Dutch-born British physician and scientist, first publishing about it in 1779.

Eukaryote

Eukaryote

Eukaryota, whose members are known as eukaryotes, is a diverse domain of organisms whose cells have a nucleus. All animals, plants, fungi, and many unicellular organisms, are eukaryotes. They belong to the group of organisms Eukaryota or Eukarya, which is one of the three domains of life. Bacteria and Archaea make up the other two domains.

Multicellular organism

Multicellular organism

A multicellular organism is an organism that consists of more than one cell, in contrast to unicellular organism.

Arthropod

Arthropod

Arthropods are invertebrate animals with an exoskeleton, a segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and cuticle made of chitin, often mineralised with calcium carbonate. The arthropod body plan consists of segments, each with a pair of appendages. Arthropods are bilaterally symmetrical and their body possesses an external skeleton. In order to keep growing, they must go through stages of moulting, a process by which they shed their exoskeleton to reveal a new one. Some species have wings. They are an extremely diverse group, with up to 10 million species.

Mollusca

Mollusca

Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks. Around 85,000 extant species of molluscs are recognized. The number of fossil species is estimated between 60,000 and 100,000 additional species. The proportion of undescribed species is very high. Many taxa remain poorly studied.

Dinosaur

Dinosaur

Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 245 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is a subject of active research. They became the dominant terrestrial vertebrates after the Triassic–Jurassic extinction event 201.3 mya and their dominance continued throughout the Jurassic and Cretaceous periods. The fossil record shows that birds are feathered dinosaurs, having evolved from earlier theropods during the Late Jurassic epoch, and are the only dinosaur lineage known to have survived the Cretaceous–Paleogene extinction event approximately 66 mya. Dinosaurs can therefore be divided into avian dinosaurs—birds—and the extinct non-avian dinosaurs, which are all dinosaurs other than birds.

Mammal

Mammal

A mammal is a vertebrate animal of the class Mammalia. Mammals are characterized by the presence of milk-producing mammary glands for feeding their young, a neocortex region of the brain, fur or hair, and three middle ear bones. These characteristics distinguish them from reptiles and birds, which they diverged from in the Carboniferous Period over 300 million years ago. Around 6,400 extant species of mammals have been described and divided into 29 orders.

Bird

Bird

Birds are a group of warm-blooded vertebrates constituting the class Aves, characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweight skeleton. Birds live worldwide and range in size from the 5.5 cm (2.2 in) bee hummingbird to the 2.8 m common ostrich. There are about ten thousand living species, more than half of which are passerine, or "perching" birds. Birds have wings whose development varies according to species; the only known groups without wings are the extinct moa and elephant birds. Wings, which are modified forelimbs, gave birds the ability to fly, although further evolution has led to the loss of flight in some birds, including ratites, penguins, and diverse endemic island species. The digestive and respiratory systems of birds are also uniquely adapted for flight. Some bird species of aquatic environments, particularly seabirds and some waterbirds, have further evolved for swimming.

Hadean

Hadean

The Hadean is a geologic eon of Earth history preceding the Archean. On Earth, the Hadean coincides with the planet's formation about 4.54 billion years ago The start of the Hadean is now defined as Ma set by the age of the oldest solid material in the Solar System, found in some meteorites, about 4.567 billion years old). The Hadean ended, as defined by the International Commission on Stratigraphy (ICS), 4 billion years ago.

Archean

Archean

The Archean Eon, in older sources sometimes called the Archaeozoic, is the second of four geologic eons of Earth's history and by definition representing the time from 4 to 2.5 billion years ago. The Archean was preceded by the Hadean Eon and followed by the Proterozoic.

Age of Earth

Age of Earth

The age of Earth is estimated to be 4.54 ± 0.05 billion years (4.54 × 109 years ± 1%). This age may represent the age of Earth's accretion, or core formation, or of the material from which Earth formed. This dating is based on evidence from radiometric age-dating of meteorite material and is consistent with the radiometric ages of the oldest-known terrestrial material and lunar samples.

Subdivisions

Subdivisions of the Silurian period
Epoch Age Start
(mya)
Etymology of
Epochs and Stages
Notes
Llandovery Rhuddanian 443.8 Cefn-Rhuddan Farm, Llandovery in Carmarthenshire, Wales
Aeronian 440.8 Cwm Coed-Aeron Farm, Wales Trefawr Track near the farm is the site of the GSSP
Telychian 438.5 Pen-lan-Telych Farm, Llandovery, Wales
Wenlock Sheinwoodian 433.4 Sheinwood village, Much Wenlock and Wenlock Edge, Shropshire, England During the Wenlock, the oldest-known tracheophytes of the genus Cooksonia, appear. The complexity of slightly later Gondwana plants like Baragwanathia, which resembled a modern clubmoss, indicates a much longer history for vascular plants, extending into the early Silurian or even Ordovician. The first terrestrial animals also appear in the Wenlock, represented by air-breathing millipedes from Scotland.[23]
Homerian 430.5 Homer, Shropshire, England
Ludlow Gorstian 427.4 Gorsty village near Ludlow, Shropshire, England
Ludfordian 425.6 Ludford, Shropshire, England
Přídolí 423.0 Named after a locality at the Homolka a Přídolí nature reserve near the Prague suburb of Slivenec, Czech Republic. Přídolí is the old name of a cadastral field area.[24]

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Llandovery Epoch

Llandovery Epoch

In the geological timescale, the Llandovery Epoch occurred at the beginning of the Silurian Period. The Llandoverian Epoch follows the massive Ordovician-Silurian extinction events, which led to a large decrease in biodiversity and an opening up of ecosystems.

Rhuddanian

Rhuddanian

In the geologic timescale, the Rhuddanian is the first age of the Silurian Period and of the Llandovery Epoch. The Silurian is in the Paleozoic Era of the Phanerozoic Eon. The Rhuddanian Age began 443.8 ± 1.5 Ma and ended 440.8 ± 1.2 Ma. It succeeds the Himantian Age and precedes the Aeronian Age.

Llandovery

Llandovery

Llandovery is a market town and community in Carmarthenshire, Wales. It lies on the River Tywi and at the junction of the A40 and A483 roads, about 25 miles (40 km) north-east of Carmarthen, 27 miles (43 km) north of Swansea and 21 miles (34 km) west of Brecon.

Carmarthenshire

Carmarthenshire

Carmarthenshire is a county in the south-west of Wales. The three largest towns are Llanelli, Carmarthen and Ammanford. Carmarthen is the county town and administrative centre. The county is known as the "Garden of Wales" and is also home to the National Botanic Garden of Wales.

Aeronian

Aeronian

In the geologic timescale, the Aeronian is an age of the Llandovery Epoch of the Silurian Period of the Paleozoic Era of the Phanerozoic Eon that began 440.8 ± 1.2 Ma and ended 438.5 ± 1.1 Ma. The Aeronian Age succeeds the Rhuddanian Age and precedes the Telychian Age, all in the same epoch.

Sheinwoodian

Sheinwoodian

In the geologic timescale, the Sheinwoodian is the age of the Wenlock Epoch of the Silurian Period of the Paleozoic Era of the Phanerozoic Eon that is comprehended between 433.4 ± 0.8 Ma and 430.5 ± 0.7 Ma, approximately. The Sheinwoodian Age succeeds the Telychian Age and precedes the Homerian Age.

Much Wenlock

Much Wenlock

Much Wenlock is a market town and parish in Shropshire, England, situated on the A458 road between Shrewsbury and Bridgnorth. Nearby, to the northeast, is the Ironbridge Gorge, and the new town of Telford. The civil parish includes the villages of Homer, Wyke, Atterley, Stretton Westwood and Bourton. The population of the civil parish, according to the 2001 census, was 2,605, increasing to 2,877 at the 2011 Census. Notable historic attractions in the town are Wenlock Priory and the Guildhall.

Cooksonia

Cooksonia

Cooksonia is an extinct group of primitive land plants, treated as a genus, although probably not monophyletic. The earliest Cooksonia date from the middle of the Silurian ; the group continued to be an important component of the flora until the end of the Early Devonian, a total time span of 433 to 393 million years ago. While Cooksonia fossils are distributed globally, most type specimens come from Britain, where they were first discovered in 1937. Cooksonia includes the oldest known plant to have a stem with vascular tissue and is thus a transitional form between the primitive non-vascular bryophytes and the vascular plants.

Gondwana

Gondwana

Gondwana was a large landmass, often referred to as a supercontinent, that formed during the late Neoproterozoic and began to break up during the Jurassic period. The final stages of break-up, involving the separation of Antarctica from South America and Australia, occurred during the Paleogene. Gondwana was not considered a supercontinent by the earliest definition, since the landmasses of Baltica, Laurentia, and Siberia were separated from it. To differentiate it from the Indian region of the same name, it is also commonly called Gondwanaland.

Baragwanathia

Baragwanathia

Baragwanathia is a genus of extinct lycopsid plants of Late Silurian to Early Devonian age, fossils of which have been found in Australia, Canada, China and Czechia. The name derives from William Baragwanath who discovered the first specimens of the type species, Baragwanathia longifolia, at Thomson River.

Ordovician

Ordovician

The Ordovician is a geologic period and system, the second of six periods of the Paleozoic Era. The Ordovician spans 41.6 million years from the end of the Cambrian Period 485.4 million years ago (Mya) to the start of the Silurian Period 443.8 Mya.

Scotland

Scotland

Scotland is a country that is part of the United Kingdom. Covering the northern third of the island of Great Britain, mainland Scotland has a 96-mile (154-kilometre) border with England to the southeast and is otherwise surrounded by the Atlantic Ocean to the north and west, the North Sea to the northeast and east, and the Irish Sea to the south. It also contains more than 790 islands, principally in the archipelagos of the Hebrides and the Northern Isles. Most of the population, including the capital Edinburgh, is concentrated in the Central Belt—the plain between the Scottish Highlands and the Southern Uplands—in the Scottish Lowlands.

Paleogeography

Ordovician-Silurian boundary on Hovedøya, Norway, showing brownish late Ordovician mudstone and later dark deep-water Silurian shale. The layers have been overturned by the Caledonian orogeny.
Ordovician-Silurian boundary on Hovedøya, Norway, showing brownish late Ordovician mudstone and later dark deep-water Silurian shale. The layers have been overturned by the Caledonian orogeny.

With the supercontinent Gondwana covering the equator and much of the southern hemisphere, a large ocean occupied most of the northern half of the globe.[25] The high sea levels of the Silurian and the relatively flat land (with few significant mountain belts) resulted in a number of island chains, and thus a rich diversity of environmental settings.[25]

During the Silurian, Gondwana continued a slow southward drift to high southern latitudes, but there is evidence that the Silurian icecaps were less extensive than those of the late-Ordovician glaciation. The southern continents remained united during this period. The melting of icecaps and glaciers contributed to a rise in sea level, recognizable from the fact that Silurian sediments overlie eroded Ordovician sediments, forming an unconformity. The continents of Avalonia, Baltica, and Laurentia drifted together near the equator, starting the formation of a second supercontinent known as Euramerica.

When the proto-Europe collided with North America, the collision folded coastal sediments that had been accumulating since the Cambrian off the east coast of North America and the west coast of Europe. This event is the Caledonian orogeny, a spate of mountain building that stretched from New York State through conjoined Europe and Greenland to Norway. At the end of the Silurian, sea levels dropped again, leaving telltale basins of evaporites extending from Michigan to West Virginia, and the new mountain ranges were rapidly eroded. The Teays River, flowing into the shallow mid-continental sea, eroded Ordovician Period strata, forming deposits of Silurian strata in northern Ohio and Indiana.

The vast ocean of Panthalassa covered most of the northern hemisphere. Other minor oceans include two phases of the Tethys, the Proto-Tethys and Paleo-Tethys, the Rheic Ocean, the Iapetus Ocean (a narrow seaway between Avalonia and Laurentia), and the newly formed Ural Ocean.

Fossils of the late Silurian sea bed
Fossils of the late Silurian sea bed

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Hovedøya

Hovedøya

Hovedøya is one of several small islands off the coast of Oslo, Norway in the Oslofjord. The island is quite small, no more than 800 metres across in any direction, the total area is 0,4 square kilometre. It is well known for its lush and green nature, with a wide variety of trees, bushes and flowers. For many, many years there was a military base on the island.

Mudstone

Mudstone

Mudstone, a type of mudrock, is a fine-grained sedimentary rock whose original constituents were clays or muds. Mudstone is distinguished from shale by its lack of fissility.

Caledonian orogeny

Caledonian orogeny

The Caledonian orogeny was a mountain-building era recorded in the northern parts of the British Isles, the Scandinavian Mountains, Svalbard, eastern Greenland and parts of north-central Europe. The Caledonian orogeny encompasses events that occurred from the Ordovician to Early Devonian, roughly 490–390 million years ago (Ma). It was caused by the closure of the Iapetus Ocean when the continents and terranes of Laurentia, Baltica and Avalonia collided.

Gondwana

Gondwana

Gondwana was a large landmass, often referred to as a supercontinent, that formed during the late Neoproterozoic and began to break up during the Jurassic period. The final stages of break-up, involving the separation of Antarctica from South America and Australia, occurred during the Paleogene. Gondwana was not considered a supercontinent by the earliest definition, since the landmasses of Baltica, Laurentia, and Siberia were separated from it. To differentiate it from the Indian region of the same name, it is also commonly called Gondwanaland.

Glacier

Glacier

A glacier is a persistent body of dense ice that is constantly moving under its own weight. A glacier forms where the accumulation of snow exceeds its ablation over many years, often centuries. It acquires distinguishing features, such as crevasses and seracs, as it slowly flows and deforms under stresses induced by its weight. As it moves, it abrades rock and debris from its substrate to create landforms such as cirques, moraines, or fjords. Although a glacier may flow into a body of water, it forms only on land and is distinct from the much thinner sea ice and lake ice that form on the surface of bodies of water.

Avalonia

Avalonia

Avalonia was a microcontinent in the Paleozoic era. Crustal fragments of this former microcontinent underlie south-west Great Britain, southern Ireland, and the eastern coast of North America. It is the source of many of the older rocks of Western Europe, Atlantic Canada, and parts of the coastal United States. Avalonia is named for the Avalon Peninsula in Newfoundland.

Baltica

Baltica

Baltica is a paleocontinent that formed in the Paleoproterozoic and now constitutes northwestern Eurasia, or Europe north of the Trans-European Suture Zone and west of the Ural Mountains. The thick core of Baltica, the East European Craton, is more than three billion years old and formed part of the Rodinia supercontinent at c. 1 Ga.

Laurentia

Laurentia

Laurentia or the North American Craton is a large continental craton that forms the ancient geological core of North America. Many times in its past, Laurentia has been a separate continent, as it is now in the form of North America, although originally it also included the cratonic areas of Greenland and also the northwestern part of Scotland, known as the Hebridean Terrane. During other times in its past, Laurentia has been part of larger continents and supercontinents and itself consists of many smaller terranes assembled on a network of Early Proterozoic orogenic belts. Small microcontinents and oceanic islands collided with and sutured onto the ever-growing Laurentia, and together formed the stable Precambrian craton seen today.

Continental drift

Continental drift

Continental drift is the hypothesis that the Earth's continents have moved over geologic time relative to each other, thus appearing to have "drifted" across the ocean bed. The idea of continental drift has been subsumed into the science of plate tectonics, which studies the movement of the continents as they ride on plates of the Earth's lithosphere.

Equator

Equator

The equator is a circle of latitude that divides a spheroid, such as Earth, into the northern and southern hemispheres. On Earth, it is an imaginary line located at 0 degrees latitude, about 40,075 km (24,901 mi) in circumference, halfway between the North and South poles. The term can also be used for any other celestial body that is roughly spherical.

New York (state)

New York (state)

New York, often called New York state, is a state in the Northeastern United States. With 20.2 million people enumerated at the 2020 United States census, its highest decennial count ever, it is the fourth-most populous state in the United States as of 2021. Approximately 44% of the state's population lives in New York City, including 25% in the boroughs of Brooklyn and Queens; and 15% of the state's population is on the remainder of Long Island, the most populous island in the United States. With a total area of 54,556 square miles (141,300 km2), New York is the 27th-largest U.S. state by area. The state is bordered by New Jersey and Pennsylvania to its south, and Connecticut, Massachusetts, and Vermont to its east; it shares a maritime border with Rhode Island, east of Long Island; and an international border with the Canadian provinces of Quebec to its north and Ontario to its northwest.

Evaporite

Evaporite

An evaporite is a water-soluble sedimentary mineral deposit that results from concentration and crystallization by evaporation from an aqueous solution. There are two types of evaporite deposits: marine, which can also be described as ocean deposits, and non-marine, which are found in standing bodies of water such as lakes. Evaporites are considered sedimentary rocks and are formed by chemical sediments.

Climate and sea level

The Silurian period was once believed to have enjoyed relatively stable and warm temperatures, in contrast with the extreme glaciations of the Ordovician before it and the extreme heat of the ensuing Devonian; however, it is now known that the global climate underwent many drastic fluctuations throughout the Silurian,[26][27] evidenced by numerous major carbon and oxygen isotope excursions during this geologic period.[28][29][30] Sea levels rose from their Hirnantian low throughout the first half of the Silurian; they subsequently fell throughout the rest of the period, although smaller scale patterns are superimposed on this general trend; fifteen high-stands (periods when sea levels were above the edge of the continental shelf) can be identified, and the highest Silurian sea level was probably around 140 metres (459 ft) higher than the lowest level reached.[25]

During this period, the Earth entered a warm greenhouse phase, supported by high CO2 levels of 4500 ppm, and warm shallow seas covered much of the equatorial land masses.[31] Early in the Silurian, glaciers retreated back into the South Pole until they almost disappeared in the middle of Silurian.[27] Layers of broken shells (called coquina) provide strong evidence of a climate dominated by violent storms generated then as now by warm sea surfaces.[32]

Perturbations

The climate and carbon cycle appear to be rather unsettled during the Silurian, which had a higher frequency of isotopic excursions (indicative of climate fluctuations) than any other period.[25] The Ireviken event, Mulde event and Lau event each represent isotopic excursions following a minor mass extinction[33] and associated with rapid sea-level change. Each one leaves a similar signature in the geological record, both geochemically and biologically; pelagic (free-swimming) organisms were particularly hard hit, as were brachiopods, corals and trilobites, and extinctions rarely occur in a rapid series of fast bursts.[25][30] The climate fluctuations are best explained by a sequence of glaciations, but the lack of tillites in the middle to late Silurian make this explanation problematic.[34]

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Hirnantian

Hirnantian

The Hirnantian is the final internationally recognized stage of the Ordovician Period of the Paleozoic Era. It was of short duration, lasting about 1.4 million years, from 445.2 to 443.8 Ma. The early part of the Hirnantian was characterized by cold temperatures, major glaciation, and a severe drop in sea level. In the latter part of the Hirnantian, temperatures rose, the glaciers melted, and sea level returned to the same or to a slightly higher level than it had been prior to the glaciation.

Earth

Earth

Earth is the third planet from the Sun and the only place known in the universe where life has originated and found habitability. While Earth may not contain the largest volumes of water in the Solar System, only Earth sustains liquid surface water, extending over 70.8% of the Earth with its ocean, making Earth an ocean world. Earth's polar regions currently retain most of all other water with large sheets of ice covering ocean and land, dwarfing Earth's groundwater, lakes, rivers and atmospheric water. Land, consisting of continents and islands, extends over 29.2% of the Earth and is widely covered by vegetation. Below Earth's surface material lies Earth's crust consisting of several slowly moving tectonic plates, which interact to produce mountain ranges, volcanoes, and earthquakes. Earth's liquid outer core generates a magnetic field that shapes the magnetosphere of Earth, largely deflecting destructive solar winds and cosmic radiation.

Greenhouse

Greenhouse

A greenhouse is a structure with walls and roof made chiefly of transparent material, such as glass, in which plants requiring regulated climatic conditions are grown.These structures range in size from small sheds to industrial-sized buildings. A miniature greenhouse is known as a cold frame. The interior of a greenhouse exposed to sunlight becomes significantly warmer than the external temperature, protecting its contents in cold weather.

Glacier

Glacier

A glacier is a persistent body of dense ice that is constantly moving under its own weight. A glacier forms where the accumulation of snow exceeds its ablation over many years, often centuries. It acquires distinguishing features, such as crevasses and seracs, as it slowly flows and deforms under stresses induced by its weight. As it moves, it abrades rock and debris from its substrate to create landforms such as cirques, moraines, or fjords. Although a glacier may flow into a body of water, it forms only on land and is distinct from the much thinner sea ice and lake ice that form on the surface of bodies of water.

South Pole

South Pole

The South Pole, also known as the Geographic South Pole, Terrestrial South Pole or 90th Parallel South, is the southernmost point on Earth and lies antipodally on the opposite side of Earth from the North Pole, at a distance of 12,430 miles in all directions. It is one of the two points where Earth's axis of rotation intersects its surface.

Coquina

Coquina

Coquina is a sedimentary rock that is composed either wholly or almost entirely of the transported, abraded, and mechanically sorted fragments of the shells of mollusks, trilobites, brachiopods, or other invertebrates. The term coquina comes from the Spanish word for "cockle" and "shellfish".

Carbon cycle

Carbon cycle

The carbon cycle is the biogeochemical cycle by which carbon is exchanged among the biosphere, pedosphere, geosphere, hydrosphere, and atmosphere of Earth. Carbon is the main component of biological compounds as well as a major component of many minerals such as limestone. Along with the nitrogen cycle and the water cycle, the carbon cycle comprises a sequence of events that are key to make Earth capable of sustaining life. It describes the movement of carbon as it is recycled and reused throughout the biosphere, as well as long-term processes of carbon sequestration to and release from carbon sinks. Carbon sinks in the land and the ocean each currently take up about one-quarter of anthropogenic carbon emissions each year.

Ireviken event

Ireviken event

The Ireviken event was the first of three relatively minor extinction events during the Silurian period. It occurred at the Llandovery/Wenlock boundary. The event is best recorded at Ireviken, Gotland, where over 50% of trilobite species became extinct; 80% of the global conodont species also became extinct in this interval.

Mulde event

Mulde event

The Mulde event was an anoxic event, and marked the second of three relatively minor mass extinctions during the Silurian period. It coincided with a global drop in sea level, and is closely followed by an excursion in geochemical isotopes. Its onset is synchronous with the deposition of the Fröel Formation in Gotland. Perceived extinction in the conodont fauna, however, likely represent a change in the depositional environment of sedimentary sequences rather than a genuine biological extinction.

Lau event

Lau event

The Lau event was the last of three relatively minor mass extinctions during the Silurian period. It had a major effect on the conodont fauna, but barely scathed the graptolites, though they suffered an extinction very shortly thereafter termed the Kozlowskii event that some authors have suggested was coeval with the Lau event and only appears asynchronous due to taphonomic reasons. It coincided with a global low point in sea level caused by glacioeustasy and is closely followed by an excursion in geochemical isotopes in the ensuing late Ludfordian faunal stage and a change in depositional regime.

Flora and fauna

The Silurian period has been viewed by some palaeontologists as an extended recovery interval following the Late Ordovician mass extinction, which interrupted the cascading increase in biodiversity that had continuously gone on throughout the Cambrian and most of the Ordovician.[35]

The Silurian was the first period to see megafossils of extensive terrestrial biota in the form of moss-like miniature forests along lakes and streams and networks of large, mycorrhizal nematophytes, heralding the beginning of the Silurian-Devonian Terrestrial Revolution.[12][13][36] However, the land fauna did not have a major impact on the Earth until it diversified in the Devonian.[25]

The first fossil records of vascular plants, that is, land plants with tissues that carry water and food, appeared in the second half of the Silurian Period.[37] The earliest-known representatives of this group are Cooksonia. Most of the sediments containing Cooksonia are marine in nature. Preferred habitats were likely along rivers and streams. Baragwanathia appears to be almost as old, dating to the early Ludlow (420 million years) and has branching stems and needle-like leaves of 10–20 centimetres (3.9–7.9 in). The plant shows a high degree of development in relation to the age of its fossil remains. Fossils of this plant have been recorded in Australia,[38][39] Canada,[40] and China.[41] Eohostimella heathana is an early, probably terrestrial, "plant" known from compression fossils[42] of Early Silurian (Llandovery) age.[43] The chemistry of its fossils is similar to that of fossilised vascular plants, rather than algae.[42]

The earliest-known animals fully adapted to terrestrial conditions appear during the Mid-Silurian, including the millipede Pneumodesmus.[23] Some evidence also suggests the presence of predatory trigonotarbid arachnoids and myriapods in Late Silurian facies.[15] Predatory invertebrates would indicate that simple food webs were in place that included non-predatory prey animals. Extrapolating back from Early Devonian biota, Andrew Jeram et al. in 1990[44] suggested a food web based on as-yet-undiscovered detritivores and grazers on micro-organisms.[45]

The first bony fish, the Osteichthyes, appeared, represented by the Acanthodians covered with bony scales. Fish reached considerable diversity and developed movable jaws, adapted from the supports of the front two or three gill arches. A diverse fauna of eurypterids (sea scorpions)—some of them several meters in length—prowled the shallow Silurian seas of North America; many of their fossils have been found in New York state. Leeches also made their appearance during the Silurian Period. Brachiopods, bryozoa, molluscs, hederelloids, tentaculitoids, crinoids and trilobites were abundant and diverse.[46] Endobiotic symbionts were common in the corals and stromatoporoids.[47][48]

Reef abundance was patchy; sometimes, fossils are frequent, but at other points, are virtually absent from the rock record.[25]

Discover more about Flora and fauna related topics

Moss

Moss

Mosses are small, non-vascular flowerless plants in the taxonomic division Bryophyta sensu stricto. Bryophyta may also refer to the parent group bryophytes, which comprise liverworts, mosses, and hornworts. Mosses typically form dense green clumps or mats, often in damp or shady locations. The individual plants are usually composed of simple leaves that are generally only one cell thick, attached to a stem that may be branched or unbranched and has only a limited role in conducting water and nutrients. Although some species have conducting tissues, these are generally poorly developed and structurally different from similar tissue found in vascular plants. Mosses do not have seeds and after fertilisation develop sporophytes with unbranched stalks topped with single capsules containing spores. They are typically 0.2–10 cm (0.1–3.9 in) tall, though some species are much larger. Dawsonia, the tallest moss in the world, can grow to 50 cm (20 in) in height. There are approximately 12,000 species.

Cooksonia

Cooksonia

Cooksonia is an extinct group of primitive land plants, treated as a genus, although probably not monophyletic. The earliest Cooksonia date from the middle of the Silurian ; the group continued to be an important component of the flora until the end of the Early Devonian, a total time span of 433 to 393 million years ago. While Cooksonia fossils are distributed globally, most type specimens come from Britain, where they were first discovered in 1937. Cooksonia includes the oldest known plant to have a stem with vascular tissue and is thus a transitional form between the primitive non-vascular bryophytes and the vascular plants.

Baragwanathia

Baragwanathia

Baragwanathia is a genus of extinct lycopsid plants of Late Silurian to Early Devonian age, fossils of which have been found in Australia, Canada, China and Czechia. The name derives from William Baragwanath who discovered the first specimens of the type species, Baragwanathia longifolia, at Thomson River.

Eohostimella

Eohostimella

Eohostimella heathana is an early, probably terrestrial, "plant" known from compression fossils of Early Silurian age. The chemistry of its fossils is similar to that of fossilised vascular plants, rather than algae. Its anatomy constitutes upright, cylindrical tubes, with a thickened outer cortex, which might have contained traces of lignin or a similar compound, even though no tracheids or similar vessels have been found; the lignin-like compound was presumably associated with its thick outer cortex. It branched dichotomously and might have borne small spines. It was probably affiliated with the rhyniophytes.

Invertebrate

Invertebrate

Invertebrates are a paraphyletic group of animals that neither possess nor develop a vertebral column, derived from the notochord. This is a grouping including all animals apart from the chordate subphylum Vertebrata. Familiar examples of invertebrates include arthropods, mollusks, annelids, echinoderms and cnidarians.

Food web

Food web

A food web is the natural interconnection of food chains and a graphical representation of what-eats-what in an ecological community. Another name for food web is consumer-resource system. Ecologists can broadly lump all life forms into one of two categories called trophic levels: 1) the autotrophs, and 2) the heterotrophs. To maintain their bodies, grow, develop, and to reproduce, autotrophs produce organic matter from inorganic substances, including both minerals and gases such as carbon dioxide. These chemical reactions require energy, which mainly comes from the Sun and largely by photosynthesis, although a very small amount comes from bioelectrogenesis in wetlands, and mineral electron donors in hydrothermal vents and hot springs. These trophic levels are not binary, but form a gradient that includes complete autotrophs, which obtain their sole source of carbon from the atmosphere, mixotrophs, which are autotrophic organisms that partially obtain organic matter from sources other than the atmosphere, and complete heterotrophs that must feed to obtain organic matter.

Detritivore

Detritivore

Detritivores are heterotrophs that obtain nutrients by consuming detritus. There are many kinds of invertebrates, vertebrates and plants that carry out coprophagy. By doing so, all these detritivores contribute to decomposition and the nutrient cycles. They should be distinguished from other decomposers, such as many species of bacteria, fungi and protists, which are unable to ingest discrete lumps of matter, but instead live by absorbing and metabolizing on a molecular scale. The terms detritivore and decomposer are often used interchangeably, but they describe different organisms. Detritivores are usually arthropods and help in the process of remineralization. Detritivores perform the first stage of remineralization, by fragmenting the dead plant matter, allowing decomposers to perform the second stage of remineralization.

Acanthodii

Acanthodii

Acanthodii or acanthodians is an extinct class of gnathostomes. They are currently considered to represent a paraphyletic grade of various fish lineages basal to extant Chondrichthyes, which includes living sharks, rays, and chimaeras. Acanthodians possess a mosaic of features shared with both osteichthyans and chondrichthyans. In general body shape, they were similar to modern sharks, but their epidermis was covered with tiny rhomboid platelets like the scales of holosteians.

Jaw

Jaw

The jaw is any opposable articulated structure at the entrance of the mouth, typically used for grasping and manipulating food. The term jaws is also broadly applied to the whole of the structures constituting the vault of the mouth and serving to open and close it and is part of the body plan of humans and most animals.

Gill

Gill

A gill is a respiratory organ that many aquatic organisms use to extract dissolved oxygen from water and to excrete carbon dioxide. The gills of some species, such as hermit crabs, have adapted to allow respiration on land provided they are kept moist. The microscopic structure of a gill presents a large surface area to the external environment. Branchia is the zoologists' name for gills.

Eurypterid

Eurypterid

Eurypterids, often informally called sea scorpions, are a group of extinct arthropods that form the order Eurypterida. The earliest known eurypterids date to the Darriwilian stage of the Ordovician period 467.3 million years ago. The group is likely to have appeared first either during the Early Ordovician or Late Cambrian period. With approximately 250 species, the Eurypterida is the most diverse Paleozoic chelicerate order. Following their appearance during the Ordovician, eurypterids became major components of marine faunas during the Silurian, from which the majority of eurypterid species have been described. The Silurian genus Eurypterus accounts for more than 90% of all known eurypterid specimens. Though the group continued to diversify during the subsequent Devonian period, the eurypterids were heavily affected by the Late Devonian extinction event. They declined in numbers and diversity until becoming extinct during the Permian–Triassic extinction event 251.9 million years ago.

Fossil

Fossil

A fossil is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved in amber, hair, petrified wood and DNA remnants. The totality of fossils is known as the fossil record.

Source: "Silurian", Wikipedia, Wikimedia Foundation, (2023, March 11th), https://en.wikipedia.org/wiki/Silurian.

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Notes
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References
  • Emiliani, Cesare. (1992). Planet Earth : Cosmology, Geology, & the Evolution of Life & the Environment. Cambridge University Press. (Paperback Edition ISBN 0-521-40949-7)
  • Mikulic, DG, DEG Briggs, and J Kluessendorf. 1985. A new exceptionally preserved biota from the Lower Silurian of Wisconsin, USA. Philosophical Transactions of the Royal Society of London, 311B:75-86.
  • Moore, RA; Briggs, DEG; Braddy, SJ; Anderson, LI; Mikulic, DG; Kluessendorf, J (2005). "A new synziphosurine (Chelicerata: Xiphosura) from the Late Llandovery (Silurian) Waukesha Lagerstatte, Wisconsin, USA". Journal of Paleontology. 79 (2): 242–250. doi:10.1666/0022-3360(2005)0792.0.co;2. S2CID 56570105.
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