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Ordovician

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Ordovician
485.4 ± 1.9 – 443.8 ± 1.5 Ma
Chronology
Etymology
Name formalityFormal
Name ratified1960
Usage information
Celestial bodyEarth
Regional usageGlobal (ICS)
Time scale(s) usedICS Time Scale
Definition
Chronological unitPeriod
Stratigraphic unitSystem
First proposed byCharles Lapworth, 1879
Time span formalityFormal
Lower boundary definitionFAD of the Conodont Iapetognathus fluctivagus
Lower boundary GSSPGreenpoint section, Green Point, Newfoundland, Canada
49°40′58″N 57°57′55″W / 49.6829°N 57.9653°W / 49.6829; -57.9653
Lower GSSP ratified2000[5]
Upper boundary definitionFAD of the Graptolite Akidograptus ascensus
Upper boundary GSSPDob's Linn, Moffat, U.K.
55°26′24″N 3°16′12″W / 55.4400°N 3.2700°W / 55.4400; -3.2700
Upper GSSP ratified1984[6][7]
Atmospheric and climatic data
Sea level above present day180 m; rising to 220 m in Caradoc and falling sharply to 140 m in end-Ordovician glaciations[8]

The Ordovician (/ɔːrdəˈvɪʃi.ən, -d-, -ˈvɪʃən/ or-də-VISH-ee-ən, -⁠doh-, -⁠VISH-ən)[9] is a geologic period and system, the second of six periods of the Paleozoic Era. The Ordovician spans 41.6 million years from the end of the Cambrian Period 485.4 million years ago (Mya) to the start of the Silurian Period 443.8 Mya.[10]

The Ordovician, named after the Welsh tribe of the Ordovices, was defined by Charles Lapworth in 1879 to resolve a dispute between followers of Adam Sedgwick and Roderick Murchison, who were placing the same rock beds in North Wales in the Cambrian and Silurian systems, respectively.[11] Lapworth recognized that the fossil fauna in the disputed strata were different from those of either the Cambrian or the Silurian systems, and placed them in a system of their own. The Ordovician received international approval in 1960 (forty years after Lapworth's death), when it was adopted as an official period of the Paleozoic Era by the International Geological Congress.

Life continued to flourish during the Ordovician as it did in the earlier Cambrian Period, although the end of the period was marked by the Ordovician–Silurian extinction events. Invertebrates, namely molluscs and arthropods, dominated the oceans, with members of the latter group probably starting their establishment on land during this time, becoming fully established by the Devonian. The first land plants are known from this period. The Great Ordovician Biodiversification Event considerably increased the diversity of life. Fish, the world's first true vertebrates, continued to evolve, and those with jaws may have first appeared late in the period. About 100 times as many meteorites struck the Earth per year during the Ordovician compared with today.[12]

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Cambrian

Cambrian

The Cambrian Period is the first geological period of the Paleozoic Era, and of the Phanerozoic Eon. The Cambrian lasted 53.4 million years from the end of the preceding Ediacaran Period 538.8 million years ago (mya) to the beginning of the Ordovician Period 485.4 mya. Its subdivisions, and its base, are somewhat in flux. The period was established as "Cambrian series" by Adam Sedgwick, who named it after Cambria, the Latin name for 'Cymru' (Wales), where Britain's Cambrian rocks are best exposed. Sedgwick identified the layer as part of his task, along with Roderick Murchison, to subdivide the large "Transition Series", although the two geologists disagreed for a while on the appropriate categorization. The Cambrian is unique in its unusually high proportion of lagerstätte sedimentary deposits, sites of exceptional preservation where "soft" parts of organisms are preserved as well as their more resistant shells. As a result, our understanding of the Cambrian biology surpasses that of some later periods.

Celtic Britons

Celtic Britons

The Britons, also known as Celtic Britons or Ancient Britons, were the people of Celtic language and culture who inhabited Great Britain from at least the British Iron Age until the High Middle Ages, at which point they diverged into the Welsh, Cornish and Bretons. They spoke Common Brittonic, the ancestor of the modern Brittonic languages.

Charles Lapworth

Charles Lapworth

Charles Lapworth FRS FGS was a headteacher and an English geologist who pioneered faunal analysis using index fossils and identified the Ordovician period.

Adam Sedgwick

Adam Sedgwick

Adam Sedgwick was a British geologist and Anglican priest, one of the founders of modern geology. He proposed the Cambrian and Devonian period of the geological timescale. Based on work which he did on Welsh rock strata, he proposed the Cambrian period in 1835, in a joint publication in which Roderick Murchison also proposed the Silurian period. Later in 1840, to resolve what later became known as the Great Devonian Controversy about rocks near the boundary between the Silurian and Carboniferous periods, he and Murchison proposed the Devonian period.

Fossil

Fossil

A fossil is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved in amber, hair, petrified wood and DNA remnants. The totality of fossils is known as the fossil record.

Fauna

Fauna

Fauna is all of the animal life present in a particular region or time. The corresponding term for plants is flora, and for fungi, it is funga. Flora, fauna, funga and other forms of life are collectively referred to as biota. Zoologists and paleontologists use fauna to refer to a typical collection of animals found in a specific time or place, e.g. the "Sonoran Desert fauna" or the "Burgess Shale fauna". Paleontologists sometimes refer to a sequence of faunal stages, which is a series of rocks all containing similar fossils. The study of animals of a particular region is called faunistics.

International Union of Geological Sciences

International Union of Geological Sciences

The International Union of Geological Sciences (IUGS) is an international non-governmental organization devoted to international cooperation in the field of geology.

Arthropod

Arthropod

Arthropods are invertebrate animals with an exoskeleton, a segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and cuticle made of chitin, often mineralised with calcium carbonate. The arthropod body plan consists of segments, each with a pair of appendages. Arthropods are bilaterally symmetrical and their body possesses an external skeleton. In order to keep growing, they must go through stages of moulting, a process by which they shed their exoskeleton to reveal a new one. Some species have wings. They are an extremely diverse group, with up to 10 million species.

Devonian

Devonian

The Devonian is a geologic period and system of the Paleozoic era, spanning 60.3 million years from the end of the Silurian, 419.2 million years ago (Mya), to the beginning of the Carboniferous, 358.9 Mya. It is named after Devon, England, where rocks from this period were first studied.

Great Ordovician Biodiversification Event

Great Ordovician Biodiversification Event

The Great Ordovician Biodiversification Event (GOBE), was an evolutionary radiation of animal life throughout the Ordovician period, 40 million years after the Cambrian explosion, whereby the distinctive Cambrian fauna fizzled out to be replaced with a Paleozoic fauna rich in suspension feeder and pelagic animals.

Fish

Fish

Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% of living fish species are ray-finned fish, belonging to the class Actinopterygii, with around 99% of those being teleosts.

Gnathostomata

Gnathostomata

Gnathostomata are the jawed vertebrates. Gnathostome diversity comprises roughly 60,000 species, which accounts for 99% of all living vertebrates, including humans. In addition to opposing jaws, living gnathostomes have true teeth, paired appendages, the elastomeric protein of elastin, and a horizontal semicircular canal of the inner ear, along with physiological and cellular anatomical characters such as the myelin sheaths of neurons, and an adaptive immune system that has the discrete lymphoid organs of spleen and thymus, and uses V(D)J recombination to create antigen recognition sites, rather than using genetic recombination in the variable lymphocyte receptor gene.

Subdivisions

A number of regional terms have been used to subdivide the Ordovician Period. In 2008, the ICS erected a formal international system of subdivisions.[13] There exist Baltoscandic, British, Siberian, North American, Australian, Chinese Mediterranean and North-Gondwanan regional stratigraphic schemes.[14]

The Ordovician Period in Britain was traditionally broken into Early (Tremadocian and Arenig), Middle (Llanvirn (subdivided into Abereiddian and Llandeilian) and Llandeilo) and Late (Caradoc and Ashgill) epochs. The corresponding rocks of the Ordovician System are referred to as coming from the Lower, Middle, or Upper part of the column. The faunal stages (subdivisions of epochs) from youngest to oldest are:

Late Ordovician

  • Hirnantian/Gamach (Ashgill)
  • Rawtheyan/Richmond (Ashgill)
  • Cautleyan/Richmond (Ashgill)
  • Pusgillian/Maysville/Richmond (Ashgill)

Middle Ordovician

  • Trenton (Caradoc)
  • Onnian/Maysville/Eden (Caradoc)
  • Actonian/Eden (Caradoc)
  • Marshbrookian/Sherman (Caradoc)
  • Longvillian/Sherman (Caradoc)
  • Soudleyan/Kirkfield (Caradoc)
  • Harnagian/Rockland (Caradoc)
  • Costonian/Black River (Caradoc)
  • Chazy (Llandeilo)
  • Llandeilo (Llandeilo)
  • Whiterock (Llanvirn)
  • Llanvirn (Llanvirn)

Early Ordovician

  • Cassinian (Arenig)
  • Arenig/Jefferson/Castleman (Arenig)
  • Tremadoc/Deming/Gaconadian (Tremadoc)

British stages

The Tremadoc corresponds to the (modern) Tremadocian. The Floian corresponds to the lower Arenig; the Arenig continues until the early Darriwilian, subsuming the Dapingian. The Llanvirn occupies the rest of the Darriwilian, and terminates with it at the base of the Late Ordovician. The Sandbian represents the first half of the Caradoc; the Caradoc ends in the mid-Katian, and the Ashgill represents the last half of the Katian, plus the Hirnantian.[15]

Ordovician regional series and stages
ICS series ICS stage British series British stage North American series North American stage Australian series Australian stage Chinese series Chinese stage
Upper Ordovician Hirnantian Ashgill Hirnantian Cincinnati Gamach Upper Ordovician Bolinda Late Ordovician Hirnantian
Katian Rawthey Richmond Chientangkiang
Cautley Maysville Easton Neichiashan
Pusgill Eden
Caradoc Strefford Mohawk Chatfield
Cheney
Sandbian Burrell Turin Gisborne
Aureluc Whiterock Chazy
Middle Ordovician Darriwilian Llanvirn Llandeilo Middle Ordovician Darriwiliane Middle Ordovician Darriwilian
Abereiddy Not defined
Dapingian Arenig Fenn Early Ordovician Yapeen Dapingian
Whitland Ranger Castlemaine
Ibex Black Hills Chewton
Bendigo
Lower Ordovician Floian Moridun Tule Lancefield Lower Ordovician Floian
Tremadocian Tremadoc Migneint Stairs Tremadocian
Cressage Skullrock

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International Classification for Standards

International Classification for Standards

International Classification for Standards (ICS) is an international classification system for technical standards. It is designed to cover every economic sector and virtually every activity of humankind where technical standards may be used.

Gondwana

Gondwana

Gondwana was a large landmass, often referred to as a supercontinent, that formed during the late Neoproterozoic and began to break up during the Jurassic period. The final stages of break-up, involving the separation of Antarctica from South America and Australia, occurred during the Paleogene. Gondwana was not considered a supercontinent by the earliest definition, since the landmasses of Baltica, Laurentia, and Siberia were separated from it. To differentiate it from the Indian region of the same name, it is also commonly called Gondwanaland.

Arenig

Arenig

In geology, the Arenig is a time interval during the Ordovician period and also the suite of rocks which were deposited during this interval.

Llandeilo Group

Llandeilo Group

In geology, the Llandeilo Group is the middle subdivision of the British Ordovician rocks. It was first described and named by Sir Roderick Murchison from the neighborhood of Llandeilo in Carmarthenshire. In the type area it consists of a series of slaty rocks, shales, calcareous flagstones and sandstones; the calcareous middle portion is sometimes termed the Llandeilo limestone; and in the upper portion volcanic rocks are intercalated.

Caradoc Series

Caradoc Series

In geology, Caradoc Series is the name introduced by Roderick Murchison in 1839 for the sandstone series of Caer Caradoc in Shropshire, England. It is the fifth of the six subdivisions of the Ordovician System, comprising all those rocks deposited worldwide during the Caradocian Age.

Stage (stratigraphy)

Stage (stratigraphy)

In chronostratigraphy, a stage is a succession of rock strata laid down in a single age on the geologic timescale, which usually represents millions of years of deposition. A given stage of rock and the corresponding age of time will by convention have the same name, and the same boundaries.

Hirnantian

Hirnantian

The Hirnantian is the final internationally recognized stage of the Ordovician Period of the Paleozoic Era. It was of short duration, lasting about 1.4 million years, from 445.2 to 443.8 Ma. The early part of the Hirnantian was characterized by cold temperatures, major glaciation, and a severe drop in sea level. In the latter part of the Hirnantian, temperatures rose, the glaciers melted, and sea level returned to the same or to a slightly higher level than it had been prior to the glaciation.

Series (stratigraphy)

Series (stratigraphy)

Series are subdivisions of rock layers based on the age of the rock and formally defined by international conventions of the geological timescale. A series is therefore a sequence of strata defining a chronostratigraphic unit. Series are subdivisions of systems and are themselves divided into stages.

Katian

Katian

The Katian is the second stage of the Upper Ordovician. It is preceded by the Sandbian and succeeded by the Hirnantian Stage. The Katian began 453 million years ago and lasted for about 7.8 million years until the beginning of the Hirnantian 445.2 million years ago. During the Katian the climate cooled which started the Late Ordovician glaciation.

Sandbian

Sandbian

The Sandbian is the first stage of the Upper Ordovician. It follows the Darriwilian and is succeeded by the Katian. Its lower boundary is defined as the first appearance datum of the graptolite species Nemagraptus gracilis around 458.4 million years ago. The Sandbian lasted for about 5.4 million years until the beginning of the Katian around 453 million years ago.

Darriwilian

Darriwilian

The Darriwilian is the upper stage of the Middle Ordovician. It is preceded by the Dapingian and succeeded by the Upper Ordovician Sandbian Stage. The lower boundary of the Darriwilian is defined as the first appearance of the graptolite species Undulograptus austrodentatus around 467.3 million years ago. It lasted for about 8.9 million years until the beginning of the Sandbian around 458.4 million years ago. This stage of the Ordovician was marked by the beginning of the Andean-Saharan glaciation.

Dapingian

Dapingian

The Dapingian is the third stage of the Ordovician period and the first stage of the Middle Ordovician series. It is preceded by the Floian and succeeded by the Darriwilian. The top of the Floian is defined as the first appearance of the conodont species Baltoniodus triangularis which happened about 470 million years ago. The Dapingian lasted for about 2.7 million years until about 467.3 million years ago.

Paleogeography and tectonics

Paleogeographic map of the Earth in the middle Ordovician, 470 million years ago
Paleogeographic map of the Earth in the middle Ordovician, 470 million years ago

During the Ordovician, the southern continents were assembled into Gondwana, which reached from north of the equator to the South Pole. The Panthalassic Ocean, centered in the northern hemisphere, covered over half the globe.[16] At the start of the period, the continents of Laurentia (in present-day North America), Siberia, and Baltica (present-day northern Europe) were separated from Gondwana by over 5,000 kilometres (3,100 mi) of ocean. These smaller continents were also sufficiently widely separated from each other to develop distinct communities of benthic organisms.[17] The small continent of Avalonia had just rifted from Gondwana and began to move north towards Baltica and Laurentia, opening the Rheic Ocean between Gondwana and Avalonia.[18][19][20] Avalonia collided with Baltica towards the end of Ordovician.[21]

Other geographic features of the Ordovician world included the Tornquist Sea, which separated Avalonia from Baltica;[17] the Aegir Ocean, which separated Baltica from Siberia;[22] and an oceanic area between Siberia, Baltica, and Gondwana which expanded to become the Paleoasian Ocean in Carboniferous time. The Mongol-Okhotsk Ocean formed a deep embayment between Siberia and the Central Mongolian terranes. Most of the terranes of central Asia were part of an equatorial archipelago whose geometry is poorly constrained by the available evidence.[23]

The period was one of extensive, widespread tectonism and volcanism. However, orogenesis (mountain-building) was not primarily due to continent-continent collisions. Instead, mountains arose along active continental margins during accretion of arc terranes or ribbon microcontinents. Accretion of new crust was limited to the Iapetus margin of Laurentia; elsewhere, the pattern was of rifting in back-arc basins followed by remerger. This reflected episodic switching from extension to compression. The initiation of new subduction reflected a global reorganization of tectonic plates centered on the amalgamation of Gondwana.[24][17]

The Taconic orogeny, a major mountain-building episode, was well under way in Cambrian times.[25] This continued into the Ordovician, when at least two volcanic island arcs collided with Laurentia to form the Appalachian Mountains. Laurentia was otherwise tectonically stable. An island arc accreted to South China during the period, while subduction along north China (Sulinheer) resulted in the emplacement of ophiolites.[26]

The ash fall of the Millburg/Big Bentonite bed, at about 454 Ma, was the largest in the last 590 million years. This had a dense rock equivalent volume of as much as 1,140 cubic kilometres (270 cu mi). Remarkably, this appears to have had little impact on life.[27]

There was vigorous tectonic activity along northwest margin of Gondwana during the Floian, 478 Ma, recorded in the Central Iberian Zone of Spain. The activity reached as far as Turkey by the end of Ordovician. The opposite margin of Gondwana, in Australia, faced a set of island arcs.[17] The accretion of these arcs to the eastern margin of Gondwana was responsible for the Benambran Orogeny of eastern Australia.[28][29] Subduction also took place along what is now Argentina (Famatinian Orogeny) at 450 Ma.[30] This involved significant back arc rifting.[17] The interior of Gondwana was tectonically quiet until the Triassic.[17]

Towards the end of the period, Gondwana began to drift across the South Pole. This contributed to the Hibernian glaciation and the associated extinction event.[31]

Ordovician meteor event

The Ordovician meteor event is a proposed shower of meteors that occurred during the Middle Ordovician Epoch, about 467.5 ± 0.28 million years ago, due to the break-up of the L chondrite parent body.[32] It is not associated with any major extinction event.[33][34][35]

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Gondwana

Gondwana

Gondwana was a large landmass, often referred to as a supercontinent, that formed during the late Neoproterozoic and began to break up during the Jurassic period. The final stages of break-up, involving the separation of Antarctica from South America and Australia, occurred during the Paleogene. Gondwana was not considered a supercontinent by the earliest definition, since the landmasses of Baltica, Laurentia, and Siberia were separated from it. To differentiate it from the Indian region of the same name, it is also commonly called Gondwanaland.

Equator

Equator

The equator is a circle of latitude that divides a spheroid, such as Earth, into the northern and southern hemispheres. On Earth, it is an imaginary line located at 0 degrees latitude, about 40,075 km (24,901 mi) in circumference, halfway between the North and South poles. The term can also be used for any other celestial body that is roughly spherical.

South Pole

South Pole

The South Pole, also known as the Geographic South Pole, Terrestrial South Pole or 90th Parallel South, is the southernmost point on Earth and lies antipodally on the opposite side of Earth from the North Pole, at a distance of 12,430 miles in all directions. It is one of the two points where Earth's axis of rotation intersects its surface.

Laurentia

Laurentia

Laurentia or the North American Craton is a large continental craton that forms the ancient geological core of North America. Many times in its past, Laurentia has been a separate continent, as it is now in the form of North America, although originally it also included the cratonic areas of Greenland and also the northwestern part of Scotland, known as the Hebridean Terrane. During other times in its past, Laurentia has been part of larger continents and supercontinents and itself consists of many smaller terranes assembled on a network of Early Proterozoic orogenic belts. Small microcontinents and oceanic islands collided with and sutured onto the ever-growing Laurentia, and together formed the stable Precambrian craton seen today.

North America

North America

North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Caribbean Sea, and to the west and south by the Pacific Ocean. Because it is on the North American Tectonic Plate, Greenland is included as a part of North America geographically.

Siberia (continent)

Siberia (continent)

Siberia, also known as Angaraland and Angarida, is an ancient craton in the heart of Siberia. Today forming the Central Siberian Plateau, it formed an independent landmass prior to its fusion into Pangea during the Late Carboniferous-Permian. The Verkhoyansk Sea, a passive continental margin, was fringing the Siberian Craton to the east in what is now the East Siberian Lowland.

Baltica

Baltica

Baltica is a paleocontinent that formed in the Paleoproterozoic and now constitutes northwestern Eurasia, or Europe north of the Trans-European Suture Zone and west of the Ural Mountains. The thick core of Baltica, the East European Craton, is more than three billion years old and formed part of the Rodinia supercontinent at c. 1 Ga.

Avalonia

Avalonia

Avalonia was a microcontinent in the Paleozoic era. Crustal fragments of this former microcontinent underlie south-west Great Britain, southern Ireland, and the eastern coast of North America. It is the source of many of the older rocks of Western Europe, Atlantic Canada, and parts of the coastal United States. Avalonia is named for the Avalon Peninsula in Newfoundland.

Rheic Ocean

Rheic Ocean

The Rheic Ocean was an ocean which separated two major palaeocontinents, Gondwana and Laurussia (Laurentia-Baltica-Avalonia). One of the principal oceans of the Palaeozoic, its sutures today stretch 10,000 km (6,200 mi) from Mexico to Turkey and its closure resulted in the assembly of the supercontinent Pangaea and the formation of the Variscan–Alleghenian–Ouachita orogenies.

Mongol-Okhotsk Ocean

Mongol-Okhotsk Ocean

The Mongol-Okhotsk Ocean also known as the Khangai-Khantey Ocean was an ancient ocean, present during the Mesozoic. Oceanic rocks from the ancient ocean floor are preserved in the Mongol-Okhotsk suture zone extending through Mongolia to the Sea of Okhotsk.

Appalachian Mountains

Appalachian Mountains

The Appalachian Mountains, often called the Appalachians, are a system of mountains in eastern to northeastern North America. The Appalachians first formed roughly 480 million years ago during the Ordovician Period. They once reached elevations similar to those of the Alps and the Rocky Mountains before experiencing natural erosion. The Appalachian chain is a barrier to east–west travel, as it forms a series of alternating ridgelines and valleys oriented in opposition to most highways and railroads running east–west.

Ordovician meteor event

Ordovician meteor event

The Ordovician meteor event was a dramatic increase in the rate at which L chondrite meteorites fell to Earth during the Middle Ordovician period, about 467.5±0.28 million years ago. This is indicated by abundant fossil L chondrite meteorites in a quarry in Sweden and enhanced concentrations of ordinary chondritic chromite grains in sedimentary rocks from this time. This temporary increase in the impact rate was most likely caused by the destruction of the L chondrite parent body 468 ± 0.3 million years ago having scattered fragments into Earth-crossing orbits, a chronology which is also supported by shock ages in numerous L chondrite meteorites that fall to Earth today. It has been speculated that this influx contributed to, or possibly even instigated, the Great Ordovician Biodiversification Event, although this has been questioned.

Geochemistry

External mold of Ordovician bivalve showing that the original aragonite shell dissolved on the sea floor, leaving a cemented mold for biological encrustation (Waynesville Formation of Franklin County, Indiana).
External mold of Ordovician bivalve showing that the original aragonite shell dissolved on the sea floor, leaving a cemented mold for biological encrustation (Waynesville Formation of Franklin County, Indiana).

The Ordovician was a time of calcite sea geochemistry in which low-magnesium calcite was the primary inorganic marine precipitate of calcium carbonate.[36] Carbonate hardgrounds were thus very common, along with calcitic ooids, calcitic cements, and invertebrate faunas with dominantly calcitic skeletons. Biogenic aragonite, like that composing the shells of most molluscs, dissolved rapidly on the sea floor after death.[37][38]

Unlike Cambrian times, when calcite production was dominated by microbial and non-biological processes, animals (and macroalgae) became a dominant source of calcareous material in Ordovician deposits.[39]

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Bivalvia

Bivalvia

Bivalvia, in previous centuries referred to as the Lamellibranchiata and Pelecypoda, is a class of marine and freshwater molluscs that have laterally compressed bodies enclosed by a shell consisting of two hinged parts. As a group, bivalves have no head and they lack some usual molluscan organs, like the radula and the odontophore. The class includes the clams, oysters, cockles, mussels, scallops, and numerous other families that live in saltwater, as well as a number of families that live in freshwater. The majority are filter feeders. The gills have evolved into ctenidia, specialised organs for feeding and breathing. Most bivalves bury themselves in sediment, where they are relatively safe from predation. Others lie on the sea floor or attach themselves to rocks or other hard surfaces. Some bivalves, such as the scallops and file shells, can swim. The shipworms bore into wood, clay, or stone and live inside these substances.

Aragonite

Aragonite

Aragonite is a carbonate mineral, one of the three most common naturally occurring crystal forms of calcium carbonate, CaCO3. It is formed by biological and physical processes, including precipitation from marine and freshwater environments.

Waynesville Formation

Waynesville Formation

The Waynesville Formation is a geologic formation in Ohio and Indiana. It preserves fossils from the Late Ordovician period.

Calcite sea

Calcite sea

A calcite sea is a sea in which low-magnesium calcite is the primary inorganic marine calcium carbonate precipitate. An aragonite sea is the alternate seawater chemistry in which aragonite and high-magnesium calcite are the primary inorganic carbonate precipitates. The Early Paleozoic and the Middle to Late Mesozoic oceans were predominantly calcite seas, whereas the Middle Paleozoic through the Early Mesozoic and the Cenozoic are characterized by aragonite seas.

Calcite

Calcite

Calcite is a carbonate mineral and the most stable polymorph of calcium carbonate (CaCO3). It is a very common mineral, particularly as a component of limestone. Calcite defines hardness 3 on the Mohs scale of mineral hardness, based on scratch hardness comparison. Large calcite crystals are used in optical equipment, and limestone composed mostly of calcite has numerous uses.

Calcium carbonate

Calcium carbonate

Calcium carbonate is a chemical compound with the chemical formula CaCO3. It is a common substance found in rocks as the minerals calcite and aragonite and is the main component of eggshells, gastropod shells, shellfish skeletons and pearls. Things containing much calcium carbonate or resembling it are described as calcareous. Calcium carbonate is the active ingredient in agricultural lime and is created when calcium ions in hard water react with carbonate ions to create limescale. It has medical use as a calcium supplement or as an antacid, but excessive consumption can be hazardous and cause hypercalcemia and digestive issues.

Carbonate hardgrounds

Carbonate hardgrounds

Carbonate hardgrounds are surfaces of synsedimentarily cemented carbonate layers that have been exposed on the seafloor. A hardground is essentially, then, a lithified seafloor. Ancient hardgrounds are found in limestone sequences and distinguished from later-lithified sediments by evidence of exposure to normal marine waters. This evidence can consist of encrusting marine organisms, borings of organisms produced through bioerosion, early marine calcite cements, or extensive surfaces mineralized by iron oxides or calcium phosphates. Modern hardgrounds are usually detected by sounding in shallow water or through remote sensing techniques like side-scan sonar.

Ooid

Ooid

Ooids are small, spheroidal, "coated" (layered) sedimentary grains, usually composed of calcium carbonate, but sometimes made up of iron- or phosphate-based minerals. Ooids usually form on the sea floor, most commonly in shallow tropical seas. After being buried under additional sediment, these ooid grains can be cemented together to form a sedimentary rock called an oolite. Oolites usually consist of calcium carbonate; these belong to the limestone rock family. Pisoids are similar to ooids, but are larger than 2 mm in diameter, often considerably larger, as with the pisoids in the hot springs at Carlsbad in the Czech Republic.

Mollusca

Mollusca

Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks. Around 85,000 extant species of molluscs are recognized. The number of fossil species is estimated between 60,000 and 100,000 additional species. The proportion of undescribed species is very high. Many taxa remain poorly studied.

Climate and sea level

The Early Ordovician climate was very hot, with intense greenhouse conditions and sea surface temperatures comparable to those during the Early Eocene Climatic Optimum.[40] By the late Early Ordovician, the Earth cooled,[41] giving way to a more temperate climate in the Middle Ordovician,[42] with the Earth likely entering the Early Palaeozoic Ice Age during the Sandbian,[43][44] and possibly as early as the Darriwilian[45] or even the Floian.[41] Evidence suggests that global temperatures rose briefly in the early Katian (Boda Event), depositing bioherms and radiating fauna across Europe.[46] Further cooling during the Hirnantian, at the end of the Ordovician, led to the Late Ordovician glaciation.[47]

The Ordovician saw the highest sea levels of the Paleozoic, and the low relief of the continents led to many shelf deposits being formed under hundreds of metres of water.[39] The sea level rose more or less continuously throughout the Early Ordovician, leveling off somewhat during the middle of the period.[39] Locally, some regressions occurred, but the sea level rise continued in the beginning of the Late Ordovician. Sea levels fell steadily due to the cooling temperatures for about 30 million years leading up to the Hirnantian glaciation. During this icy stage, sea level seems to have risen and dropped somewhat. Despite much study, the details remain unresolved.[39] In particular, some researches interpret the fluctuations in sea level as pre-Hibernian glaciation,[48] but sedimentary evidence of glaciation is lacking until the end of the period.[21] There is evidence of glaciers during the Hirnantian on the land we now know as Africa and South America, which were near the South Pole at the time, facilitating the formation of the ice caps of the Hirnantian glaciation.

As with North America and Europe, Gondwana was largely covered with shallow seas during the Ordovician. Shallow clear waters over continental shelves encouraged the growth of organisms that deposit calcium carbonates in their shells and hard parts. The Panthalassic Ocean covered much of the Northern Hemisphere, and other minor oceans included Proto-Tethys, Paleo-Tethys, Khanty Ocean, which was closed off by the Late Ordovician, Iapetus Ocean, and the new Rheic Ocean.

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Greenhouse and icehouse Earth

Greenhouse and icehouse Earth

Throughout Earth's climate history (Paleoclimate) its climate has fluctuated between two primary states: greenhouse and icehouse Earth. Both climate states last for millions of years and should not be confused with glacial and interglacial periods, which occur as alternate phases within an icehouse period and tend to last less than 1 million years. There are five known Icehouse periods in Earth's climate history, which are known as the Huronian, Cryogenian, Andean-Saharan, Late Paleozoic, and Late Cenozoic glaciations. The main factors involved in changes of the paleoclimate are believed to be the concentration of atmospheric carbon dioxide, changes in Earth's orbit, long-term changes in the solar constant, and oceanic and orogenic changes from tectonic plate dynamics. Greenhouse and icehouse periods have played key roles in the evolution of life on Earth by directly and indirectly forcing biotic adaptation and turnover at various spatial scales across time.

Late Ordovician glaciation

Late Ordovician glaciation

The Late Ordovician glaciation, also known as the Hirnantian glaciation or end-Ordovician glaciation, is the first part of the Andean-Saharan glaciation. It was centered on the Sahara region in late Ordovician, about 440–460 Ma. The major glaciation during this period is widely considered to be the leading cause of the Ordovician-Silurian extinction event. Evidence of this glaciation can be seen in places such as Morocco, Libya, South Africa, Bolivia, Argentina, and Wyoming. More evidence derived from isotopic data is that during the Late Ordovician, tropical ocean temperatures were about 5 °C cooler than present day; this would have been a major factor that aided in the glaciation process.

Glacier

Glacier

A glacier is a persistent body of dense ice that is constantly moving under its own weight. A glacier forms where the accumulation of snow exceeds its ablation over many years, often centuries. It acquires distinguishing features, such as crevasses and seracs, as it slowly flows and deforms under stresses induced by its weight. As it moves, it abrades rock and debris from its substrate to create landforms such as cirques, moraines, or fjords. Although a glacier may flow into a body of water, it forms only on land and is distinct from the much thinner sea ice and lake ice that form on the surface of bodies of water.

Gondwana

Gondwana

Gondwana was a large landmass, often referred to as a supercontinent, that formed during the late Neoproterozoic and began to break up during the Jurassic period. The final stages of break-up, involving the separation of Antarctica from South America and Australia, occurred during the Paleogene. Gondwana was not considered a supercontinent by the earliest definition, since the landmasses of Baltica, Laurentia, and Siberia were separated from it. To differentiate it from the Indian region of the same name, it is also commonly called Gondwanaland.

Ice cap

Ice cap

In glaciology, an ice cap is a mass of ice that covers less than 50,000 km2 (19,000 sq mi) of land area. Larger ice masses covering more than 50,000 km2 (19,000 sq mi) are termed ice sheets.

North America

North America

North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Caribbean Sea, and to the west and south by the Pacific Ocean. Because it is on the North American Tectonic Plate, Greenland is included as a part of North America geographically.

Europe

Europe

Europe is a continent comprising the westernmost peninsulas of Eurasia, located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It shares the continental landmass of Afro-Eurasia with both Africa and Asia. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east. Europe is commonly considered to be separated from Asia by the watershed of the Ural Mountains, the Ural River, the Caspian Sea, the Greater Caucasus, the Black Sea and the waterways of the Turkish Straits.

Northern Hemisphere

Northern Hemisphere

The Northern Hemisphere is the half of Earth that is north of the Equator. For other planets in the Solar System, north is defined as being in the same celestial hemisphere relative to the invariable plane of the solar system as Earth's North Pole.

Proto-Tethys Ocean

Proto-Tethys Ocean

The Proto-Tethys or Theic Ocean was an ancient ocean that existed from the latest Ediacaran to the Carboniferous.

Paleo-Tethys Ocean

Paleo-Tethys Ocean

The Paleo-Tethys or Palaeo-Tethys Ocean was an ocean located along the northern margin of the paleocontinent Gondwana that started to open during the Middle Cambrian, grew throughout the Paleozoic, and finally closed during the Late Triassic; existing for about 400 million years.

Khanty Ocean

Khanty Ocean

Khanty Ocean or Khanty-Mansi Ocean was a small ocean located between Baltica and an island arc near Siberia, from near the end of the Precambrian time to the Silurian. The ocean is named after two indigenous peoples in Siberia.

Iapetus Ocean

Iapetus Ocean

The Iapetus Ocean was an ocean that existed in the late Neoproterozoic and early Paleozoic eras of the geologic timescale. The Iapetus Ocean was situated in the southern hemisphere, between the paleocontinents of Laurentia, Baltica and Avalonia. The ocean disappeared with the Acadian, Caledonian and Taconic orogenies, when these three continents joined to form one big landmass called Euramerica. The "southern" Iapetus Ocean has been proposed to have closed with the Famatinian and Taconic orogenies, meaning a collision between Western Gondwana and Laurentia.

Life

A diorama depicting Ordovician flora and fauna
A diorama depicting Ordovician flora and fauna

For most of the Late Ordovician life continued to flourish, but at and near the end of the period there were mass-extinction events that seriously affected conodonts and planktonic forms like graptolites. The trilobites Agnostida and Ptychopariida completely died out, and the Asaphida were much reduced. Brachiopods, bryozoans and echinoderms were also heavily affected, and the endocerid cephalopods died out completely, except for possible rare Silurian forms. The Ordovician–Silurian extinction events may have been caused by an ice age that occurred at the end of the Ordovician Period, due to the expansion of the first terrestrial plants,[49] as the end of the Late Ordovician was one of the coldest times in the last 600 million years of Earth's history.

Fauna

Endoceras, one of the largest predators of the Ordovician
Endoceras, one of the largest predators of the Ordovician
Fossiliferous limestone slab from the Liberty Formation (Upper Ordovician) of Caesar Creek State Park near Waynesville, Ohio.
Fossiliferous limestone slab from the Liberty Formation (Upper Ordovician) of Caesar Creek State Park near Waynesville, Ohio.
The trilobite Isotelus from Wisconsin
The trilobite Isotelus from Wisconsin

On the whole, the fauna that emerged in the Ordovician were the template for the remainder of the Palaeozoic. The fauna was dominated by tiered communities of suspension feeders, mainly with short food chains. The ecological system reached a new grade of complexity far beyond that of the Cambrian fauna, which has persisted until the present day.[39]

Though less famous than the Cambrian explosion, the Ordovician radiation (also known as the Great Ordovician Biodiversification Event)[17] was no less remarkable; marine faunal genera increased fourfold, resulting in 12% of all known Phanerozoic marine fauna.[50] Another change in the fauna was the strong increase in filter-feeding organisms.[51] The trilobite, inarticulate brachiopod, archaeocyathid, and eocrinoid faunas of the Cambrian were succeeded by those that dominated the rest of the Paleozoic, such as articulate brachiopods, cephalopods, and crinoids. Articulate brachiopods, in particular, largely replaced trilobites in shelf communities. Their success epitomizes the greatly increased diversity of carbonate shell-secreting organisms in the Ordovician compared to the Cambrian.[52]

Aegirocassis, a large filter-feeding hurdiid radiodont from Morocco
Aegirocassis, a large filter-feeding hurdiid radiodont from Morocco

Ordovician geography had its effect on the diversity of fauna. The widely separated continents of Laurentia and Baltica, then positioned close to the tropics and boasting many shallow seas rich in life, developed a distinct trilobite fauna from the trilobite fauna of Gondwana, and Gondwana developed distinct fauna in its tropical and temperature zones. However, tropical articulate brachiopods had a more cosmopolitan distribution, with less diversity on different continents. Faunas become less provincial later in the Ordovician, though they were still distinguishable into the late Ordovician.[53]

Pentecopterus, the earliest known eurypterid, and found in Iowa
Pentecopterus, the earliest known eurypterid, and found in Iowa

Trilobites in particular were rich and diverse. Trilobites in the Ordovician were very different from their predecessors in the Cambrian. Many trilobites developed bizarre spines and nodules to defend against predators such as primitive eurypterids and nautiloids while other trilobites such as Aeglina prisca evolved to become swimming forms. Some trilobites even developed shovel-like snouts for ploughing through muddy sea bottoms. Another unusual clade of trilobites known as the trinucleids developed a broad pitted margin around their head shields.[54] Some trilobites such as Asaphus kowalewski evolved long eyestalks to assist in detecting predators whereas other trilobite eyes in contrast disappeared completely.[55] Molecular clock analyses suggest that early arachnids started living on land by the end of the Ordovician.[56] Although solitary corals date back to at least the Cambrian, reef-forming corals appeared in the early Ordovician, including the earliest known octocorals,[57][58] corresponding to an increase in the stability of carbonate and thus a new abundance of calcifying animals.[39] Brachiopods surged in diversity, adapting to almost every type of marine environment.[59][60][61] Molluscs, which appeared during the Cambrian or even the Ediacaran, became common and varied, especially bivalves, gastropods, and nautiloid cephalopods.[62][63] Cephalopods diversified from shallow marine tropical environments to dominate almost all marine environments.[64] Graptolites, which evolved in the preceding Cambrian period, thrived in the oceans. This includes the distinctive Nemagraptus gracilis graptolite fauna, which was distributed widely during peak sea levels in the Sandbian.[65][21][21] Some new cystoids and crinoids appeared. It was long thought that the first true vertebrates (fish — Ostracoderms) appeared in the Ordovician, but recent discoveries in China reveal that they probably originated in the Early Cambrian.[66] The first gnathostome (jawed fish) may have appeared in the Late Ordovician epoch.[67] Chitinozoans, which first appeared late in the Wuliuan, exploded in diversity during the Tremadocian, quickly becoming globally widespread.[68][69] Several groups of endobiotic symbionts appeared in the Ordovician.[70][71]

In the Early Ordovician, trilobites were joined by many new types of organisms, including tabulate corals, strophomenid, rhynchonellid, and many new orthid brachiopods, bryozoans, planktonic graptolites and conodonts, and many types of molluscs and echinoderms, including the ophiuroids ("brittle stars") and the first sea stars. Nevertheless, the arthropods remained abundant; all the Late Cambrian orders continued, and were joined by the new group Phacopida. The first evidence of land plants also appeared (see evolutionary history of life).

In the Middle Ordovician, the trilobite-dominated Early Ordovician communities were replaced by generally more mixed ecosystems, in which brachiopods, bryozoans, molluscs, cornulitids, tentaculitids and echinoderms all flourished, tabulate corals diversified and the first rugose corals appeared. The planktonic graptolites remained diverse, with the Diplograptina making their appearance. One of the earliest known armoured agnathan ("ostracoderm") vertebrates, Arandaspis, dates from the Middle Ordovician.[72] During the Middle Ordovician there was a large increase in the intensity and diversity of bioeroding organisms. This is known as the Ordovician Bioerosion Revolution.[73] It is marked by a sudden abundance of hard substrate trace fossils such as Trypanites, Palaeosabella, Petroxestes and Osprioneides. Bioerosion became an important process, particularly in the thick calcitic skeletons of corals, bryozoans and brachiopods, and on the extensive carbonate hardgrounds that appear in abundance at this time.

Flora

Green algae were common in the Late Cambrian (perhaps earlier) and in the Ordovician. Terrestrial plants probably evolved from green algae, first appearing as tiny non-vascular forms resembling liverworts, in the middle to late Ordovician.[75] Fossil spores found in Ordovician sedimentary rock are typical of bryophytes.[76]

Colonization of land would have been limited to shorelines
Colonization of land would have been limited to shorelines

Among the first land fungi may have been arbuscular mycorrhiza fungi (Glomerales), playing a crucial role in facilitating the colonization of land by plants through mycorrhizal symbiosis, which makes mineral nutrients available to plant cells; such fossilized fungal hyphae and spores from the Ordovician of Wisconsin have been found with an age of about 460 million years ago, a time when the land flora most likely only consisted of plants similar to non-vascular bryophytes.[77]

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Diorama

Diorama

A diorama is a replica of a scene, typically a three-dimensional full-size or miniature model, sometimes enclosed in a glass showcase for a museum. Dioramas are often built by hobbyists as part of related hobbies such as military vehicle modeling, miniature figure modeling, or aircraft modeling.

Conodont

Conodont

Conodonts are an extinct group of agnathan (jawless) vertebrates resembling eels, classified in the class Conodonta. For many years, they were known only from their tooth-like oral elements, which are usually found in isolation and are now called conodont elements. Knowledge about soft tissues remains limited. They existed in the world's oceans for over 300 million years, from the Cambrian to the beginning of the Jurassic. Conodont elements are widely used as index fossils, fossils used to define and identify geological periods. The animals are also called Conodontophora to avoid ambiguity.

Plankton

Plankton

Plankton are the diverse collection of organisms found in water that are unable to propel themselves against a current. The individual organisms constituting plankton are called plankters. In the ocean, they provide a crucial source of food to many small and large aquatic organisms, such as bivalves, fish and whales.

Agnostida

Agnostida

Agnostida is an order of arthropod which have classically been seen as a group of highly modified trilobites, though some recent research has doubted this placement. Regardless, they appear to be close relatives as part of the Artiopoda. They are present in the Lower Cambrian fossil record along with trilobites from the Redlichiida, Corynexochida, and Ptychopariida orders, and were highly diverse throughout the Cambrian. Agnostidan diversity severely declined during the Cambrian-Ordovician transition, and the last agnostidans went extinct in the Late Ordovician.

Asaphida

Asaphida

Asaphida is a large, morphologically diverse order of trilobites found in marine strata dated from the Middle Cambrian until their extinction during the Silurian. Asaphida contains six superfamilies, but no suborders. Asaphids comprise some 20% of described fossil trilobites.

Brachiopod

Brachiopod

Brachiopods, phylum Brachiopoda, are a phylum of trochozoan animals that have hard "valves" (shells) on the upper and lower surfaces, unlike the left and right arrangement in bivalve molluscs. Brachiopod valves are hinged at the rear end, while the front can be opened for feeding or closed for protection. Two major categories are traditionally recognized, articulate and inarticulate brachiopods. The word "articulate" is used to describe the tooth-and-groove structures of the valve-hinge which is present in the articulate group, and absent from the inarticulate group. This is the leading diagnostic skeletal feature, by which the two main groups can be readily distinguished as fossils. Articulate brachiopods have toothed hinges and simple, vertically-oriented opening and closing muscles. Conversely, inarticulate brachiopods have weak, untoothed hinges and a more complex system of vertical and oblique (diagonal) muscles used to keep the two valves aligned. In many brachiopods, a stalk-like pedicle projects from an opening near the hinge of one of the valves, known as the pedicle or ventral valve. The pedicle, when present, keeps the animal anchored to the seabed but clear of sediment which would obstruct the opening.

Bryozoa

Bryozoa

Bryozoa are a phylum of simple, aquatic invertebrate animals, nearly all living in sedentary colonies. Typically about 0.5 millimetres long, they have a special feeding structure called a lophophore, a "crown" of tentacles used for filter feeding. Most marine bryozoans live in tropical waters, but a few are found in oceanic trenches and polar waters. The bryozoans are classified as the marine bryozoans (Stenolaemata), freshwater bryozoans (Phylactolaemata), and mostly-marine bryozoans (Gymnolaemata), a few members of which prefer brackish water. 5,869 living species are known. At least two genera are solitary ; the rest are colonial.

Echinoderm

Echinoderm

An echinoderm is any member of the phylum Echinodermata. The adults are recognisable by their radial symmetry, and include starfish, brittle stars, sea urchins, sand dollars, and sea cucumbers, as well as the sea lilies or "stone lilies". Adult echinoderms are found on the sea bed at every ocean depth, from the intertidal zone to the abyssal zone. The phylum contains about 7,000 living species, making it the second-largest grouping of deuterostomes, after the chordates. Echinoderms are the largest entirely marine phylum. The first definitive echinoderms appeared near the start of the Cambrian.

Cephalopod

Cephalopod

A cephalopod is any member of the molluscan class Cephalopoda such as a squid, octopus, cuttlefish, or nautilus. These exclusively marine animals are characterized by bilateral body symmetry, a prominent head, and a set of arms or tentacles modified from the primitive molluscan foot. Fishers sometimes call cephalopods "inkfish", referring to their common ability to squirt ink. The study of cephalopods is a branch of malacology known as teuthology.

Endoceras

Endoceras

Endoceras is an extinct genus of large, straight shelled cephalopods that gives its name to the Nautiloid order Endocerida. The genus lived during the middle and upper Ordovician 485 to 419 million years ago. The cross section in the mature portion is slightly wider than high, but is narrower laterally in the young. Sutures are straight and transverse. Endoceras has a large siphuncle, located close to the ventral margin, composed of concave segments, especially in the young but which may be tubular in the adult stage. Endocones are simple, subcircular in cross section, and penetrated by a narrow tube which may contain diaphragms reminiscent of the Ellesmerocerid ancestor.

Isotelus

Isotelus

Isotelus is a genus of asaphid trilobites from the middle and upper Ordovician period, fairly common in the Northeastern United States, northwest Manitoba, southwestern Quebec and southeastern Ontario. One species, Isotelus rex, is currently the world's largest trilobite ever found as a complete fossil, and was probably exceeded in size only by Hungioides bohemicus, in which the specimens probably exceeding Isotelus rex in size are only known from partial remains.

Cambrian explosion

Cambrian explosion

The Cambrian explosion, Cambrian radiation, Cambrian diversification, or the Biological Big Bang refers to an interval of time approximately 538.8 million years ago in the Cambrian Period when practically all major animal phyla started appearing in the fossil record. It lasted for about 13 – 25 million years and resulted in the divergence of most modern metazoan phyla. The event was accompanied by major diversification in other groups of organisms as well.

End of the period

The Ordovician came to a close in a series of extinction events that, taken together, comprise the second largest of the five major extinction events in Earth's history in terms of percentage of genera that became extinct. The only larger one was the Permian–Triassic extinction event.

The extinctions occurred approximately 447–444 million years ago and mark the boundary between the Ordovician and the following Silurian Period. At that time all complex multicellular organisms lived in the sea, and about 49% of genera of fauna disappeared forever; brachiopods and bryozoans were greatly reduced, along with many trilobite, conodont and graptolite families.

The most commonly accepted theory is that these events were triggered by the onset of cold conditions in the late Katian, followed by an ice age, in the Hirnantian faunal stage, that ended the long, stable greenhouse conditions typical of the Ordovician.

The ice age was possibly not long-lasting. Oxygen isotopes in fossil brachiopods show its duration may have been only 0.5 to 1.5 million years.[78] Other researchers (Page et al.) estimate more temperate conditions did not return until the late Silurian.

The late Ordovician glaciation event was preceded by a fall in atmospheric carbon dioxide (from 7000 ppm to 4400 ppm).[79][80] The dip may have been caused by a burst of volcanic activity that deposited new silicate rocks, which draw CO2 out of the air as they erode.[80] Another possibility is that bryophytes and lichens, which colonized land in the middle to late Ordovician, may have increased weathering enough to draw down CO2 levels.[75] The drop in CO2 selectively affected the shallow seas where most organisms lived. As the southern supercontinent Gondwana drifted over the South Pole, ice caps formed on it, which have been detected in Upper Ordovician rock strata of North Africa and then-adjacent northeastern South America, which were south-polar locations at the time.

As glaciers grew, the sea level dropped, and the vast shallow intra-continental Ordovician seas withdrew, which eliminated many ecological niches. When they returned, they carried diminished founder populations that lacked many whole families of organisms. They then withdrew again with the next pulse of glaciation, eliminating biological diversity with each change.[81] Species limited to a single epicontinental sea on a given landmass were severely affected.[38] Tropical lifeforms were hit particularly hard in the first wave of extinction, while cool-water species were hit worst in the second pulse.[38]

Those species able to adapt to the changing conditions survived to fill the ecological niches left by the extinctions. For example, there is evidence the oceans became more deeply oxygenated during the glaciation, allowing unusual benthic organisms (Hirnantian fauna) to colonize the depths. These organisms were cosmopolitan in distribution and present at most latitudes.[53]

At the end of the second event, melting glaciers caused the sea level to rise and stabilise once more. The rebound of life's diversity with the permanent re-flooding of continental shelves at the onset of the Silurian saw increased biodiversity within the surviving Orders. Recovery was characterized by an unusual number of "Lazarus taxa", disappearing during the extinction and reappearing well into the Silurian, which suggests that the taxa survived in small numbers in refugia.[82]

An alternate extinction hypothesis suggested that a ten-second gamma-ray burst could have destroyed the ozone layer and exposed terrestrial and marine surface-dwelling life to deadly ultraviolet radiation and initiated global cooling.[83]

Recent work considering the sequence stratigraphy of the Late Ordovician argues that the mass extinction was a single protracted episode lasting several hundred thousand years, with abrupt changes in water depth and sedimentation rate producing two pulses of last occurrences of species.[84]

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Extinction event

Extinction event

An extinction event is a widespread and rapid decrease in the biodiversity on Earth. Such an event is identified by a sharp change in the diversity and abundance of multicellular organisms. It occurs when the rate of extinction increases with respect to the background extinction rate and the rate of speciation. Estimates of the number of major mass extinctions in the last 540 million years range from as few as five to more than twenty. These differences stem from disagreement as to what constitutes a "major" extinction event, and the data chosen to measure past diversity.

History of Earth

History of Earth

The history of Earth concerns the development of planet Earth from its formation to the present day. Nearly all branches of natural science have contributed to understanding of the main events of Earth's past, characterized by constant geological change and biological evolution.

Genus

Genus

Genus is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nomenclature, the genus name forms the first part of the binomial species name for each species within the genus.E.g. Panthera leo (lion) and Panthera onca (jaguar) are two species within the genus Panthera. Panthera is a genus within the family Felidae.

Permian–Triassic extinction event

Permian–Triassic extinction event

The Permian–Triassic extinction event, also known as the End-Permian extinction event and colloquially as the Great Dying, forms the boundary between the Permian and Triassic geologic periods, and with them the Paleozoic and Mesozoic eras respectively, approximately 251.9 million years ago. As the largest of the "Big Five" mass extinctions of the Phanerozoic, it is the Earth's most severe known extinction event, with the extinction of 57% of biological families, 83% of genera, 81% of marine species and 70% of terrestrial vertebrate species. It is the largest known mass extinction of insects. There is evidence for one to three distinct pulses, or phases, of extinction.

Conodont

Conodont

Conodonts are an extinct group of agnathan (jawless) vertebrates resembling eels, classified in the class Conodonta. For many years, they were known only from their tooth-like oral elements, which are usually found in isolation and are now called conodont elements. Knowledge about soft tissues remains limited. They existed in the world's oceans for over 300 million years, from the Cambrian to the beginning of the Jurassic. Conodont elements are widely used as index fossils, fossils used to define and identify geological periods. The animals are also called Conodontophora to avoid ambiguity.

Ice age

Ice age

An ice age is a long period of reduction in the temperature of Earth's surface and atmosphere, resulting in the presence or expansion of continental and polar ice sheets and alpine glaciers. Earth's climate alternates between ice ages and greenhouse periods, during which there are no glaciers on the planet. Earth is in the Quaternary glaciation. Individual pulses of cold climate within an ice age are termed glacial periods, and intermittent warm periods within an ice age are called interglacials or interstadials.

Greenhouse

Greenhouse

A greenhouse is a structure with walls and roof made chiefly of transparent material, such as glass, in which plants requiring regulated climatic conditions are grown.These structures range in size from small sheds to industrial-sized buildings. A miniature greenhouse is known as a cold frame. The interior of a greenhouse exposed to sunlight becomes significantly warmer than the external temperature, protecting its contents in cold weather.

Late Ordovician glaciation

Late Ordovician glaciation

The Late Ordovician glaciation, also known as the Hirnantian glaciation or end-Ordovician glaciation, is the first part of the Andean-Saharan glaciation. It was centered on the Sahara region in late Ordovician, about 440–460 Ma. The major glaciation during this period is widely considered to be the leading cause of the Ordovician-Silurian extinction event. Evidence of this glaciation can be seen in places such as Morocco, Libya, South Africa, Bolivia, Argentina, and Wyoming. More evidence derived from isotopic data is that during the Late Ordovician, tropical ocean temperatures were about 5 °C cooler than present day; this would have been a major factor that aided in the glaciation process.

Bryophyte

Bryophyte

The Bryophyta s.l. are a proposed taxonomic division containing three groups of non-vascular land plants (embryophytes): the liverworts, hornworts and mosses. Bryophyta s.s. consists of the mosses only. They are characteristically limited in size and prefer moist habitats although they can survive in drier environments. The bryophytes consist of about 20,000 plant species. Bryophytes produce enclosed reproductive structures, but they do not produce flowers or seeds. They reproduce sexually by spores and asexually by fragmentation or the production of gemmae. Though bryophytes were considered a paraphyletic group in recent years, almost all of the most recent phylogenetic evidence supports the monophyly of this group, as originally classified by Wilhelm Schimper in 1879. The term bryophyte comes from Ancient Greek βρύον (brúon) 'tree moss, liverwort', and φυτόν (phutón) 'plant'.

Gondwana

Gondwana

Gondwana was a large landmass, often referred to as a supercontinent, that formed during the late Neoproterozoic and began to break up during the Jurassic period. The final stages of break-up, involving the separation of Antarctica from South America and Australia, occurred during the Paleogene. Gondwana was not considered a supercontinent by the earliest definition, since the landmasses of Baltica, Laurentia, and Siberia were separated from it. To differentiate it from the Indian region of the same name, it is also commonly called Gondwanaland.

North Africa

North Africa

North Africa, or Northern Africa, is a region encompassing the northern portion of the African continent. There is no singularly accepted scope for the region, and it is sometimes defined as stretching from the Atlantic shores of Mauritania in the west, to Egypt's Suez Canal in the east.

Gamma-ray burst

Gamma-ray burst

In gamma-ray astronomy, gamma-ray bursts (GRBs) are immensely energetic explosions that have been observed in distant galaxies. They are the most energetic and luminous electromagnetic events since the Big Bang. Bursts can last from ten milliseconds to several hours. After an initial flash of gamma rays, a longer-lived "afterglow" is usually emitted at longer wavelengths.

Source: "Ordovician", Wikipedia, Wikimedia Foundation, (2023, March 19th), https://en.wikipedia.org/wiki/Ordovician.

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