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Moulting

From Wikipedia, in a visual modern way
A dragonfly in its radical final moult, metamorphosing from an aquatic nymph to a winged adult.
A dragonfly in its radical final moult, metamorphosing from an aquatic nymph to a winged adult.

In biology, moulting (British English), or molting (American English), also known as sloughing, shedding, or in many invertebrates, ecdysis, is the manner in which an animal routinely casts off a part of its body (often, but not always, an outer layer or covering), either at specific times of the year, or at specific points in its life cycle.

In medieval times it was also known as "mewing" (from the French verb "muer", to moult),[1] a term that lives on in the name of Britain's Royal Mews where the King's hawks used to be kept during moulting time before becoming horse stables after Tudor times.

Moulting can involve shedding the epidermis (skin), pelage (hair, feathers, fur, wool), or other external layer. In some groups, other body parts may be shed, for example, the entire exoskeleton in arthropods, including the wings in some insects.

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Biology

Biology

Biology is the scientific study of life. It is a natural science with a broad scope but has several unifying themes that tie it together as a single, coherent field. For instance, all organisms are made up of cells that process hereditary information encoded in genes, which can be transmitted to future generations. Another major theme is evolution, which explains the unity and diversity of life. Energy processing is also important to life as it allows organisms to move, grow, and reproduce. Finally, all organisms are able to regulate their own internal environments.

British English

British English

British English is, according to Oxford Dictionaries, "English as used in Great Britain, as distinct from that used elsewhere". More narrowly, it can refer specifically to the English language in England, or, more broadly, to the collective dialects of English throughout the British Isles taken as a single umbrella variety, for instance additionally incorporating Scottish English, Welsh English, and Northern Irish English. Tom McArthur in the Oxford Guide to World English acknowledges that British English shares "all the ambiguities and tensions [with] the word 'British' and as a result can be used and interpreted in two ways, more broadly or more narrowly, within a range of blurring and ambiguity".

American English

American English

American English, sometimes called United States English or U.S. English, is the set of varieties of the English language native to the United States. English is the most widely spoken language in the United States and in most circumstances is the de facto common language used in government, education and commerce. Since the 20th century, American English has become the most influential form of English worldwide.

Ecdysis

Ecdysis

Ecdysis is the moulting of the cuticle in many invertebrates of the clade Ecdysozoa. Since the cuticle of these animals typically forms a largely inelastic exoskeleton, it is shed during growth and a new, larger covering is formed. The remnants of the old, empty exoskeleton are called exuviae.

Royal Mews

Royal Mews

The Royal Mews is a mews, or collection of equestrian stables, of the British royal family. In London these stables and stable-hands' quarters have occupied two main sites in turn, being located at first on the north side of Charing Cross, and then within the grounds of Buckingham Palace.

Hair

Hair

Hair is a protein filament that grows from follicles found in the dermis. Hair is one of the defining characteristics of mammals. The human body, apart from areas of glabrous skin, is covered in follicles which produce thick terminal and fine vellus hair. Most common interest in hair is focused on hair growth, hair types, and hair care, but hair is also an important biomaterial primarily composed of protein, notably alpha-keratin.

Feather

Feather

Feathers are epidermal growths that form a distinctive outer covering, or plumage, on both avian (bird) and some non-avian dinosaurs and other archosaurs. They are the most complex integumentary structures found in vertebrates and a premier example of a complex evolutionary novelty. They are among the characteristics that distinguish the extant birds from other living groups.

Fur

Fur

Fur is a thick growth of hair that covers the skin of mammals. It consists of a combination of oily guard hair on top and thick underfur beneath. The guard hair keeps moisture from reaching the skin; the underfur acts as an insulating blanket that keeps the animal warm.

Exoskeleton

Exoskeleton

An exoskeleton is an external skeleton that supports and protects an animal's body, in contrast to an internal skeleton (endoskeleton) in for example, a human. In usage, some of the larger kinds of exoskeletons are known as "shells". Examples of exoskeletons within animals include the arthropod exoskeleton shared by chelicerates, myriapods, crustaceans, and insects, as well as the shell of certain sponges and the mollusc shell shared by snails, clams, tusk shells, chitons and nautilus. Some animals, such as the turtle, have both an endoskeleton and an exoskeleton.

Arthropod

Arthropod

Arthropods are invertebrate animals with an exoskeleton, a segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and cuticle made of chitin, often mineralised with calcium carbonate. The arthropod body plan consists of segments, each with a pair of appendages. Arthropods are bilaterally symmetrical and their body possesses an external skeleton. In order to keep growing, they must go through stages of moulting, a process by which they shed their exoskeleton to reveal a new one. Some species have wings. They are an extremely diverse group, with up to 10 million species.

Wing

Wing

A wing is a type of fin that produces lift while moving through air or some other fluid. Accordingly, wings have streamlined cross-sections that are subject to aerodynamic forces and act as airfoils. A wing's aerodynamic efficiency is expressed as its lift-to-drag ratio. The lift a wing generates at a given speed and angle of attack can be one to two orders of magnitude greater than the total drag on the wing. A high lift-to-drag ratio requires a significantly smaller thrust to propel the wings through the air at sufficient lift.

Insect

Insect

Insects are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body, three pairs of jointed legs, compound eyes and one pair of antennae. Their blood is not totally contained in vessels; some circulates in an open cavity known as the haemocoel. Insects are the most diverse group of animals; they include more than a million described species and represent more than half of all known living organisms. The total number of extant species is estimated at between six and ten million; potentially over 90% of the animal life forms on Earth are insects. Insects may be found in nearly all environments, although only a small number of species reside in the oceans, which are dominated by another arthropod group, crustaceans, which recent research has indicated insects are nested within.

Examples

Group Item shed Timing Notes
Cats Fur Usually around spring-summer time Cats moult fur around spring-summer time to get rid of their "winter coat". Cats have thicker fur during the colder winter months to keep them warm, then around spring and summer they shed some of their fur to get a thinner coat for the warmer summer months. Some cats need brushing during moulting, since dead hairs can get trapped in the cat's fur.
Dogs and other canids Fur Semi-annually, spring and fall (autumn). Moulting or shedding in canids, as in all mammals,[2] is due to fluctuations in the amount of melatonin secreted by their pineal gland in response to seasonal sunlight variations rather than temperature variations. This seasonality in moulting is most preserved in Arctic breeds of dogs which shed twice each year whereas most other breeds moult once each year.
Chicken Feathers Usually autumn (non-commercial hens). Chickens generally stop laying eggs when their moulting begins and recommence laying when their new feathers have re-grown.
Mallards Feathers Mid-summer - early fall After the end of the breeding season, most mallards moult their flight feathers. As the brightly-coloured breeding plumage of the males leaves them vulnerable to predation, they lose it through moulting, replacing it with eclipse plumage that aids in camouflage until their flight feathers regrow, upon which they moult again and regain their breeding colours.[3]
Snakes Skin Regularly, when old skin is outgrown. Snakes rub against rough surfaces to assist removal of their shed skin.
Lizards Skin Regularly, when old skin is outgrown. Lizards, like snakes, rub against objects to help remove their shed skin and then consume the shed skin for calcium and other nutrients.
Amphibians Skin Regularly. Salamanders and frogs shed their skins regularly, then often eat it.
Hermit crabs Exoskeleton Regularly, when the carapace is outgrown. Land hermit crabs bury themselves for many weeks while they moult and then consume their exoskeleton.
Arachnids Exoskeleton Regularly, when the exoskeleton is outgrown. Arachnids moult regularly to grow, often becoming reclusive and fasting for long periods prior to a moult.
Insects Exoskeleton Regularly in larvae, when the exoskeleton is outgrown. In species with a "complete" metamorphosis, the final moult transforms the body, typically from a soft-bodied larva to a reproductive, winged and sometimes colourful adult. In mayflies, a winged subimago moults one last time to a winged adult.

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Cat

Cat

The cat is a domestic species of small carnivorous mammal. It is the only domesticated species in the family Felidae and is commonly referred to as the domestic cat or house cat to distinguish it from the wild members of the family. Cats are commonly kept as house pets but can also be farm cats or feral cats; the feral cat ranges freely and avoids human contact. Domestic cats are valued by humans for companionship and their ability to kill rodents. About 60 cat breeds are recognized by various cat registries.

Fur

Fur

Fur is a thick growth of hair that covers the skin of mammals. It consists of a combination of oily guard hair on top and thick underfur beneath. The guard hair keeps moisture from reaching the skin; the underfur acts as an insulating blanket that keeps the animal warm.

Dog

Dog

The dog is a domesticated descendant of the wolf. Also called the domestic dog, it is derived from the extinct Pleistocene wolf, and the modern wolf is the dog's nearest living relative. Dogs were the first species to be domesticated by hunter-gatherers over 15,000 years ago before the development of agriculture. Due to their long association with humans, dogs have expanded to a large number of domestic individuals and gained the ability to thrive on a starch-rich diet that would be inadequate for other canids.

Melatonin

Melatonin

Melatonin is a natural compound, specifically an indoleamine, produced by and found in different organisms including bacteria and eukaryotes. It was discovered by Aaron B. Lerner and colleagues in 1958 as a substance of the pineal gland from cow that could induce skin lightening in common frogs. It was subsequently discovered as a hormone released in the brain at night which controls the sleep–wake cycle in vertebrates.

Chicken

Chicken

The chicken is a domesticated junglefowl species, with attributes of wild species such as the grey and the Ceylon junglefowl that are originally from Southeast Asia. Rooster and cock are terms for adult male birds, and a younger male may be called a cockerel. A male that has been castrated is a capon. An adult female bird is called a hen, and a sexually immature female is called a pullet. Humans keep chickens primarily as a source of food or as pets. Traditionally they were also bred for cockfighting, which is still practiced in some places. Chickens domesticated for meat are broilers and for eggs are layers.

Feather

Feather

Feathers are epidermal growths that form a distinctive outer covering, or plumage, on both avian (bird) and some non-avian dinosaurs and other archosaurs. They are the most complex integumentary structures found in vertebrates and a premier example of a complex evolutionary novelty. They are among the characteristics that distinguish the extant birds from other living groups.

Mallard

Mallard

The mallard or wild duck is a dabbling duck that breeds throughout the temperate and subtropical Americas, Eurasia, and North Africa, and has been introduced to New Zealand, Australia, Peru, Brazil, Uruguay, Argentina, Chile, Colombia, the Falkland Islands, and South Africa. This duck belongs to the subfamily Anatinae of the waterfowl family Anatidae. Males have purple patches on their wings, while the females have mainly brown-speckled plumage. Both sexes have an area of white-bordered black or iridescent blue feathers called a speculum on their wings; males especially tend to have blue speculum feathers. The mallard is 50–65 cm (20–26 in) long, of which the body makes up around two-thirds the length. The wingspan is 81–98 cm (32–39 in) and the bill is 4.4 to 6.1 cm long. It is often slightly heavier than most other dabbling ducks, weighing 0.7–1.6 kg (1.5–3.5 lb). Mallards live in wetlands, eat water plants and small animals, and are social animals preferring to congregate in groups or flocks of varying sizes.

Snake

Snake

Snakes are elongated, limbless, carnivorous reptiles of the suborder Serpentes. Like all other squamates, snakes are ectothermic, amniote vertebrates covered in overlapping scales. Many species of snakes have skulls with several more joints than their lizard ancestors, enabling them to swallow prey much larger than their heads. To accommodate their narrow bodies, snakes' paired organs appear one in front of the other instead of side by side, and most have only one functional lung. Some species retain a pelvic girdle with a pair of vestigial claws on either side of the cloaca. Lizards have independently evolved elongate bodies without limbs or with greatly reduced limbs at least twenty-five times via convergent evolution, leading to many lineages of legless lizards. These resemble snakes, but several common groups of legless lizards have eyelids and external ears, which snakes lack, although this rule is not universal.

Lizard

Lizard

Lizards are a widespread group of squamate reptiles, with over 7,000 species, ranging across all continents except Antarctica, as well as most oceanic island chains. The group is paraphyletic since it excludes the snakes and Amphisbaenia although some lizards are more closely related to these two excluded groups than they are to other lizards. Lizards range in size from chameleons and geckos a few centimeters long to the 3-meter-long Komodo dragon.

Skin

Skin

Skin is the layer of usually soft, flexible outer tissue covering the body of a vertebrate animal, with three main functions: protection, regulation, and sensation.

Amphibian

Amphibian

Amphibians are four-limbed and ectothermic vertebrates of the class Amphibia. All living amphibians belong to the group Lissamphibia. They inhabit a wide variety of habitats, with most species living within terrestrial, fossorial, arboreal or freshwater aquatic ecosystems. Thus amphibians typically start out as larvae living in water, but some species have developed behavioural adaptations to bypass this.

Hermit crab

Hermit crab

Hermit crabs are anomuran decapod crustaceans of the superfamily Paguroidea that have adapted to occupy empty scavenged mollusc shells to protect their fragile exoskeletons. There are over 800 species of hermit crab, most of which possess an asymmetric abdomen concealed by a snug-fitting shell. Hermit crabs' soft (non-calcified) abdominal exoskeleton means they must occupy shelter produced by other organisms or risk being defenseless.

In birds

Loggerhead shrike moulting. Loggerhead shrike with normal plumage.
A loggerhead shrike in mid-moult (left) and with regular plumage (right).
A king penguin with developing replacement feathers, sometimes called pin feathers
A king penguin with developing replacement feathers, sometimes called pin feathers

In birds, moulting is the periodic replacement of feathers by shedding old feathers while producing new ones. Feathers are dead structures at maturity which are gradually abraded and need to be replaced. Adult birds moult at least once a year, although many moult twice and a few three times each year.[4] It is generally a slow process as birds rarely shed all their feathers at any one time; the bird must retain sufficient feathers to regulate its body temperature and repel moisture. The number and area of feathers that are shed varies. In some moulting periods, a bird may renew only the feathers on the head and body, shedding the wing and tail feathers during a later moulting period.[4] Some species of bird become flightless during an annual "wing moult" and must seek a protected habitat with a reliable food supply during that time. While the plumage may appear thin or uneven during the moult, the bird's general shape is maintained despite the loss of apparently many feathers; bald spots are typically signs of unrelated illnesses, such as gross injuries, parasites, feather pecking (especially in commercial poultry), or (in pet birds) feather plucking. Some birds will drop feathers, especially tail feathers, in what is called a "fright moult".[5]

The process of moulting in birds is as follows: First, the bird begins to shed some old feathers, then pin feathers grow in to replace the old feathers. As the pin feathers become full feathers, other feathers are shed. This is a cyclical process that occurs in many phases. It is usually symmetrical, with feather loss equal on each side of the body.[4] Because feathers make up 4–12% of a bird's body weight, it takes a large amount of energy to replace them. For this reason, moults often occur immediately after the breeding season, but while food is still abundant. The plumage produced during this time is called postnuptial plumage.[4] Prenuptial moulting occurs in red-collared widowbirds where the males replace their nonbreeding plumage with breeding plumage. It is thought that large birds can advance the moult of severely damaged feathers.[6]

Determining the process birds go through during moult can be useful in understanding breeding, migration and foraging strategies.[7] One non-invasive method of studying moult in birds is through using field photography.[8] The evolutionary and ecological forces driving moult can also be investigated using intrinsic markers such as stable hydrogen isotope (δ2H) analysis.[9] In some tropical birds, such as the common bulbul, breeding seasonality is weak at the population level, instead moult can show high seasonality with individuals probably under strong selection to match moult with peak environmental conditions.[10]

Forced moulting

In some countries, flocks of commercial layer hens are force-moulted to reinvigorate egg-laying. This usually involves complete withdrawal of their food and sometimes water for 7–14 days or up to 28 days under experimental conditions,[11] which presumably reflect standard farming practice in some countries. This causes a body weight loss of 25 to 35%,[12] which stimulates the hen to lose her feathers, but also reinvigorates egg-production. Some flocks may be force-moulted several times. In 2003, more than 75% of all flocks were force-moulted in the US.[13] Other methods of inducing a moult include low-density diets (e.g. grape pomace, cotton seed meal, alfalfa meal)[14] or dietary manipulation to create an imbalance of a particular nutrient(s). The most important among these include manipulation of minerals including sodium (Na), calcium (Ca), iodine (I) and zinc (Zn), with full or partially reduced dietary intakes.[15]

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Loggerhead shrike

Loggerhead shrike

The loggerhead shrike is a passerine bird in the family Laniidae. It is the only member of the shrike family endemic to North America; the related northern shrike occurs north of its range, however it is also found in Siberia. It is nicknamed the butcherbird after its carnivorous tendencies, as it consumes prey such as amphibians, insects, lizards, small mammals and small birds, and some prey end up displayed and stored at a site, for example in a tree. Due to its small size and weak talons, this predatory bird relies on impaling its prey upon thorns or barbed wire for facilitated consumption. The numbers of loggerhead shrike have significantly decreased in recent years, especially in Midwestern, New England and Mid-Atlantic areas.

King penguin

King penguin

The king penguin is the second largest species of penguin, smaller, but somewhat similar in appearance to the emperor penguin. There are two subspecies: A. p. patagonicus and A. p. halli; patagonicus is found in the South Atlantic and halli in the South Indian Ocean and at Macquarie Island.

Pin feather

Pin feather

A pin feather is a developing feather on a bird. This feather can grow as a new feather during the bird's infancy, or grow to replace one from moulting.

Bird

Bird

Birds are a group of warm-blooded vertebrates constituting the class Aves, characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweight skeleton. Birds live worldwide and range in size from the 5.5 cm (2.2 in) bee hummingbird to the 2.8 m common ostrich. There are about ten thousand living species, more than half of which are passerine, or "perching" birds. Birds have wings whose development varies according to species; the only known groups without wings are the extinct moa and elephant birds. Wings, which are modified forelimbs, gave birds the ability to fly, although further evolution has led to the loss of flight in some birds, including ratites, penguins, and diverse endemic island species. The digestive and respiratory systems of birds are also uniquely adapted for flight. Some bird species of aquatic environments, particularly seabirds and some waterbirds, have further evolved for swimming.

Feather

Feather

Feathers are epidermal growths that form a distinctive outer covering, or plumage, on both avian (bird) and some non-avian dinosaurs and other archosaurs. They are the most complex integumentary structures found in vertebrates and a premier example of a complex evolutionary novelty. They are among the characteristics that distinguish the extant birds from other living groups.

Feather pecking

Feather pecking

Feather pecking is a behavioural problem that occurs most frequently amongst domestic hens reared for egg production, although it does occur in other poultry such as pheasants, turkeys, ducks, broiler chickens and is sometimes seen in farmed ostriches. Feather pecking occurs when one bird repeatedly pecks at the feathers of another. The levels of severity may be recognized as mild and severe. Gentle feather pecking is considered to be a normal investigatory behaviour where the feathers of the recipient are hardly disturbed and therefore does not represent a problem. In severe feather pecking, however, the feathers of the recipient are grasped, pulled at and sometimes removed. This is painful for the receiving bird and can lead to trauma of the skin or bleeding, which in turn can lead to cannibalism and death.

Common bulbul

Common bulbul

The common bulbul is a member of the bulbul family of passerine birds. It is found in north-eastern, northern, western and central Africa.

Forced molting

Forced molting

Forced molting, sometimes known as induced molting, is the practice by some poultry industries of artificially provoking a flock to molt simultaneously, typically by withdrawing food for 7–14 days and sometimes also withdrawing water for an extended period. Forced molting is usually implemented when egg-production is naturally decreasing toward the end of the first egg-laying phase. During the forced molt, the birds cease producing eggs for at least two weeks, which allows the bird's reproductive tracts to regress and rejuvenate. After the molt, the hen's egg production rate usually peaks slightly lower than the previous peak, but egg quality is improved. The purpose of forced molting is therefore to increase egg production, egg quality, and profitability of flocks in their second or subsequent laying phases, by not allowing the hen's body the necessary time to rejuvenate during the natural cycle of feather replenishment.

Chicken

Chicken

The chicken is a domesticated junglefowl species, with attributes of wild species such as the grey and the Ceylon junglefowl that are originally from Southeast Asia. Rooster and cock are terms for adult male birds, and a younger male may be called a cockerel. A male that has been castrated is a capon. An adult female bird is called a hen, and a sexually immature female is called a pullet. Humans keep chickens primarily as a source of food or as pets. Traditionally they were also bred for cockfighting, which is still practiced in some places. Chickens domesticated for meat are broilers and for eggs are layers.

Alfalfa

Alfalfa

Alfalfa, also called lucerne, is a perennial flowering plant in the legume family Fabaceae. It is cultivated as an important forage crop in many countries around the world. It is used for grazing, hay, and silage, as well as a green manure and cover crop. The name alfalfa is used in North America. The name lucerne is the more commonly used name in the United Kingdom, South Africa, Australia, and New Zealand. The plant superficially resembles clover, especially while young, when trifoliate leaves comprising round leaflets predominate. Later in maturity, leaflets are elongated. It has clusters of small purple flowers followed by fruits spiralled in 2 to 3 turns containing 10–20 seeds. Alfalfa is native to warmer temperate climates. It has been cultivated as livestock fodder since at least the era of the ancient Greeks and Romans.

Calcium

Calcium

Calcium is a chemical element with the symbol Ca and atomic number 20. As an alkaline earth metal, calcium is a reactive metal that forms a dark oxide-nitride layer when exposed to air. Its physical and chemical properties are most similar to its heavier homologues strontium and barium. It is the fifth most abundant element in Earth's crust, and the third most abundant metal, after iron and aluminium. The most common calcium compound on Earth is calcium carbonate, found in limestone and the fossilised remnants of early sea life; gypsum, anhydrite, fluorite, and apatite are also sources of calcium. The name derives from Latin calx "lime", which was obtained from heating limestone.

Iodine

Iodine

Iodine is a chemical element with the symbol I and atomic number 53. The heaviest of the stable halogens, it exists as a semi-lustrous, non-metallic solid at standard conditions that melts to form a deep violet liquid at 114 °C (237 °F), and boils to a violet gas at 184 °C (363 °F). The element was discovered by the French chemist Bernard Courtois in 1811 and was named two years later by Joseph Louis Gay-Lussac, after the Ancient Greek Ιώδης 'violet-coloured'.

In reptiles

A young Mediterranean House Gecko in the process of moulting.
A young Mediterranean House Gecko in the process of moulting.
Close up view of snake's moulted skin A black-bearded gliding lizard moulting
Close up view of snake's moulted skin (left) and a black-bearded gliding lizard moulting (right).

Squamates periodically engage in moulting, as their skin is scaly. The most familiar example of moulting in such reptiles is when snakes "shed their skin". This is usually achieved by the snake rubbing its head against a hard object, such as a rock (or between two rocks) or piece of wood, causing the already stretched skin to split. At this point, the snake continues to rub its skin on objects, causing the end nearest the head to peel back on itself, until the snake is able to crawl out of its skin, effectively turning the moulted skin inside-out. This is similar to how one might remove a sock from one's foot by grabbing the open end and pulling it over itself. The snake's skin is often left in one piece after the moulting process, including the discarded brille (ocular scale), so that the moult is vital for maintaining the animal's quality of vision. The skins of lizards, in contrast, generally fall off in pieces.

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Reptile

Reptile

Reptiles, as most commonly defined, are the animals in the class Reptilia, a paraphyletic grouping comprising all sauropsids except birds. Living reptiles comprise turtles, crocodilians, squamates and rhynchocephalians (tuatara). As of March 2022, the Reptile Database includes about 11,700 species. In the traditional Linnaean classification system, birds are considered a separate class to reptiles. However, crocodilians are more closely related to birds than they are to other living reptiles, and so modern cladistic classification systems include birds within Reptilia, redefining the term as a clade. Other cladistic definitions abandon the term reptile altogether in favor of the clade Sauropsida, which refers to all amniotes more closely related to modern reptiles than to mammals. The study of the traditional reptile orders, customarily in combination with the study of modern amphibians, is called herpetology.

Snake

Snake

Snakes are elongated, limbless, carnivorous reptiles of the suborder Serpentes. Like all other squamates, snakes are ectothermic, amniote vertebrates covered in overlapping scales. Many species of snakes have skulls with several more joints than their lizard ancestors, enabling them to swallow prey much larger than their heads. To accommodate their narrow bodies, snakes' paired organs appear one in front of the other instead of side by side, and most have only one functional lung. Some species retain a pelvic girdle with a pair of vestigial claws on either side of the cloaca. Lizards have independently evolved elongate bodies without limbs or with greatly reduced limbs at least twenty-five times via convergent evolution, leading to many lineages of legless lizards. These resemble snakes, but several common groups of legless lizards have eyelids and external ears, which snakes lack, although this rule is not universal.

Skin

Skin

Skin is the layer of usually soft, flexible outer tissue covering the body of a vertebrate animal, with three main functions: protection, regulation, and sensation.

Sock

Sock

A sock is a piece of clothing worn on the feet and often covering the ankle or some part of the calf. Some types of shoes or boots are typically worn over socks. In ancient times, socks were made from leather or matted animal hair. In the late 16th century, machine-knit socks were first produced. Until the 1800s, both hand-made and machine-knit socks were manufactured, with the latter technique becoming more common in the 19th century.

Foot

Foot

The foot is an anatomical structure found in many vertebrates. It is the terminal portion of a limb which bears weight and allows locomotion. In many animals with feet, the foot is a separate organ at the terminal part of the leg made up of one or more segments or bones, generally including claws and or nails.

Brille

Brille

The brille is the layer of transparent, immovable disc-shaped skin or scale covering the eyes of some animals for protection, especially in animals without eyelids. The brille has evolved from a fusion of the upper and lower eyelids. Brille means "spectacles" or "glasses" in German, Norwegian, and Danish, as well as "shine" in French and Spanish.

Lizard

Lizard

Lizards are a widespread group of squamate reptiles, with over 7,000 species, ranging across all continents except Antarctica, as well as most oceanic island chains. The group is paraphyletic since it excludes the snakes and Amphisbaenia although some lizards are more closely related to these two excluded groups than they are to other lizards. Lizards range in size from chameleons and geckos a few centimeters long to the 3-meter-long Komodo dragon.

In arthropods

In arthropods, such as insects, arachnids and crustaceans, moulting is the shedding of the exoskeleton (which is often called its shell), typically to let the organism grow. This process is called ecdysis. It is commonly said that ecdysis is necessary because the exoskeleton is rigid and cannot grow like skin, but this is simplistic, ignoring the fact that most Arthropoda with soft, flexible skins also undergo ecdysis. Among other things, ecdysis permits metamorphosis, the sometimes radical difference between the morphology of successive instars, and the fact that a new skin can replace structures, such as by providing new external lenses for eyes. The new exoskeleton is initially soft but hardens after the moulting of the old exoskeleton. The old exoskeleton is called an exuviae. While moulting, insects can't breathe.[16]

Moulting phase of a southern hawker
Moulting phase of a southern hawker

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Ecdysis

Ecdysis

Ecdysis is the moulting of the cuticle in many invertebrates of the clade Ecdysozoa. Since the cuticle of these animals typically forms a largely inelastic exoskeleton, it is shed during growth and a new, larger covering is formed. The remnants of the old, empty exoskeleton are called exuviae.

Metamorphosis

Metamorphosis

Metamorphosis is a biological process by which an animal physically develops including birth transformation or hatching, involving a conspicuous and relatively abrupt change in the animal's body structure through cell growth and differentiation. Some insects, fish, amphibians, mollusks, crustaceans, cnidarians, echinoderms, and tunicates undergo metamorphosis, which is often accompanied by a change of nutrition source or behavior. Animals can be divided into species that undergo complete metamorphosis ("holometaboly"), incomplete metamorphosis ("hemimetaboly"), or no metamorphosis ("ametaboly").

Arthropod

Arthropod

Arthropods are invertebrate animals with an exoskeleton, a segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and cuticle made of chitin, often mineralised with calcium carbonate. The arthropod body plan consists of segments, each with a pair of appendages. Arthropods are bilaterally symmetrical and their body possesses an external skeleton. In order to keep growing, they must go through stages of moulting, a process by which they shed their exoskeleton to reveal a new one. Some species have wings. They are an extremely diverse group, with up to 10 million species.

Insect

Insect

Insects are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body, three pairs of jointed legs, compound eyes and one pair of antennae. Their blood is not totally contained in vessels; some circulates in an open cavity known as the haemocoel. Insects are the most diverse group of animals; they include more than a million described species and represent more than half of all known living organisms. The total number of extant species is estimated at between six and ten million; potentially over 90% of the animal life forms on Earth are insects. Insects may be found in nearly all environments, although only a small number of species reside in the oceans, which are dominated by another arthropod group, crustaceans, which recent research has indicated insects are nested within.

Arachnid

Arachnid

Arachnida is a class of joint-legged invertebrate animals (arthropods), in the subphylum Chelicerata. Arachnida includes, among others, spiders, scorpions, ticks, mites, pseudoscorpions, harvestmen, camel spiders, whip spiders and vinegaroons.

Crustacean

Crustacean

Crustaceans belong to the subphylum Crustacea,, and form a large, diverse group of arthropods including decapods, seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, amphipods and mantis shrimp. The crustacean group can be treated as a subphylum under the clade Mandibulata. It is now well accepted that the hexapods emerged deep in the Crustacean group, with the completed group referred to as Pancrustacea. Some crustaceans are more closely related to insects and the other hexapods than they are to certain other crustaceans.

Exoskeleton

Exoskeleton

An exoskeleton is an external skeleton that supports and protects an animal's body, in contrast to an internal skeleton (endoskeleton) in for example, a human. In usage, some of the larger kinds of exoskeletons are known as "shells". Examples of exoskeletons within animals include the arthropod exoskeleton shared by chelicerates, myriapods, crustaceans, and insects, as well as the shell of certain sponges and the mollusc shell shared by snails, clams, tusk shells, chitons and nautilus. Some animals, such as the turtle, have both an endoskeleton and an exoskeleton.

Skin

Skin

Skin is the layer of usually soft, flexible outer tissue covering the body of a vertebrate animal, with three main functions: protection, regulation, and sensation.

Instar

Instar

An instar is a developmental stage of arthropods, such as insects, between each moult (ecdysis), until sexual maturity is reached. Arthropods must shed the exoskeleton in order to grow or assume a new form. Differences between instars can often be seen in altered body proportions, colors, patterns, changes in the number of body segments or head width. After shedding their exoskeleton (moulting), the juvenile arthropods continue in their life cycle until they either pupate or moult again. The instar period of growth is fixed; however, in some insects, like the salvinia stem-borer moth, the number of instars depends on early larval nutrition. Some arthropods can continue to moult after sexual maturity, but the stages between these subsequent moults are generally not called instars.

Exuviae

Exuviae

In biology, exuviae are the remains of an exoskeleton and related structures that are left after ecdysozoans have moulted. The exuviae of an animal can be important to biologists as they can often be used to identify the species of the animal and even its sex.

Southern hawker

Southern hawker

The southern hawker or blue hawker is a species of hawker dragonfly.

In dogs

Most dogs moult twice each year, in the spring and autumn, depending on the breed, environment and temperature. Dogs shedding much more than usual are known as "blow coats" or "blowing coats".[17][18]

In amphibians

Both frogs and salamanders moult regularly and consume the skin, with some species moulting in pieces and others in one piece.[19]

Gallery

Discover more about Gallery related topics

Yellow-eyed penguin

Yellow-eyed penguin

The yellow-eyed penguin, known also as hoiho or tarakaka, is a species of penguin endemic to New Zealand.

Leopard frog

Leopard frog

Leopard frog is a generic name used to refer to various species in the true frog genus Lithobates. They all have similar coloration: brown or green with spots that form a leopard pattern. They are distinguished by their distribution and behavioral, morphological, and genetic differences. The range of the various species of leopard frogs extends from the Hudson Bay in Canada, throughout the United States, throughout Mexico and other parts of Central America, and possibly the very northern section of South America.

Extatosoma tiaratum

Extatosoma tiaratum

Extatosoma tiaratum, commonly known as the spiny leaf insect, the giant prickly stick insect, Macleay's spectre, or the Australian walking stick, is a large species of Australian stick insect endemic to Australia. The species has the Phasmid Study Group number PSG9.

European green lizard

European green lizard

The European green lizard is a large lizard distributed across European midlatitudes from Slovenia and eastern Austria to as far east as the Black Sea coasts of Ukraine and Turkey. It is often seen sunning on rocks or lawns, or sheltering amongst bushes.

Caelifera

Caelifera

The Caelifera are a suborder of orthopteran insects. They include the grasshoppers and grasshopper-like insects, as well as other superfamilies classified with them: the ground-hoppers (Tetrigoidea) and pygmy mole crickets (Tridactyloidea). The latter should not be confused with the mole crickets (Gryllotalpidae), which belong to the other Orthopteran sub-order Ensifera.

Tiliqua rugosa

Tiliqua rugosa

Tiliqua rugosa, most commonly known as the shingleback skink or bobtail lizard, is a short-tailed, slow-moving species of blue-tongued skink endemic to Australia. It is commonly known as the shingleback or sleepy lizard. Three of its four recognised subspecies are found in Western Australia, where the bobtail name is most frequently used. The fourth subspecies, T. rugosa asper, is the only one native to eastern Australia, where it goes by the common name of the eastern shingleback.

Source: "Moulting", Wikipedia, Wikimedia Foundation, (2022, December 10th), https://en.wikipedia.org/wiki/Moulting.

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References
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