Late Cretaceous

The Turonian is, in the ICS' geologic timescale, the second age in the Late Cretaceous Epoch, or a stage in the Upper Cretaceous Series. It spans the time between 93.9 ± 0.8 Ma and 89.8 ± 1 Ma. The Turonian is preceded by the Cenomanian Stage and underlies the Coniacian Stage.

Plate tectonics is the generally accepted scientific theory that considers the Earth's lithosphere to comprise a number of large tectonic plates which have been slowly moving since about 3.4 billion years ago. The model builds on the concept of continental drift, an idea developed during the first decades of the 20th century. Plate tectonics came to be generally accepted by geoscientists after seafloor spreading was validated in the mid to late 1960s.

The Western Interior Seaway was a large inland sea that split the continent of North America into two landmasses. The ancient sea, which existed from the early Late Cretaceous to the earliest Paleocene, connected the Gulf of Mexico, through the United States and Canada, to the Arctic Ocean. The two land masses it created were Laramidia to the west and Appalachia to the east. At its largest extent, it was 2,500 feet (760 m) deep, 600 miles (970 km) wide and over 2,000 miles (3,200 km) long.

During most of the Late Cretaceous the eastern half of North America formed Appalachia, an island land mass separated from Laramidia to the west by the Western Interior Seaway. This seaway had split North America into two massive landmasses due to a multitude of factors such as tectonism and sea-level fluctuations for nearly 40 million years. The seaway eventually expanded, divided across the Dakotas, and by the end of the Cretaceous, it retreated towards the Gulf of Mexico and the Hudson Bay. This left the island masses joined in the continent of North America as the Rocky Mountains rose. From the Cenomanian to the end of the Campanian ages of the Late Cretaceous, Appalachia was separated from the rest of North America. As the Western Interior Seaway retreated in the Maastrichtian, Laramidia and Appalachia eventually connected. Because of this, its fauna was isolated, and developed very differently from the tyrannosaur, ceratopsian, hadrosaurid, pachycephalosaur and ankylosaurid dominated fauna of the western part of North America, known as "Laramidia".

Laramidia was an island continent that existed during the Late Cretaceous period, when the Western Interior Seaway split the continent of North America in two. In the Mesozoic era, Laramidia was an island land mass separated from Appalachia to the east by the Western Interior Seaway. The seaway eventually shrank, split across the Dakotas, and retreated toward the Gulf of Mexico and the Hudson Bay. The masses joined, forming the continent of North America.

Antarctica is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean, it contains the geographic South Pole. Antarctica is the fifth-largest continent, being about 40% larger than Europe, and has an area of 14,200,000 km2 (5,500,000 sq mi). Most of Antarctica is covered by the Antarctic ice sheet, with an average thickness of 1.9 km (1.2 mi).

Insular dwarfism, a form of phyletic dwarfism, is the process and condition of large animals evolving or having a reduced body size when their population's range is limited to a small environment, primarily islands. This natural process is distinct from the intentional creation of dwarf breeds, called dwarfing. This process has occurred many times throughout evolutionary history, with examples including dinosaurs, like Europasaurus and Magyarosaurus dacus, and modern animals such as elephants and their relatives. This process, and other "island genetics" artifacts, can occur not only on islands, but also in other situations where an ecosystem is isolated from external resources and breeding. This can include caves, desert oases, isolated valleys and isolated mountains. Insular dwarfism is one aspect of the more general "island effect" or "Foster's rule", which posits that when mainland animals colonize islands, small species tend to evolve larger bodies, and large species tend to evolve smaller bodies. This is itself one aspect of island syndrome, which describes the differences in morphology, ecology, physiology and behaviour of insular species compared to their continental counterparts.

Hadrosaurids, or duck-billed dinosaurs, are members of the ornithischian family Hadrosauridae. This group is known as the duck-billed dinosaurs for the flat duck-bill appearance of the bones in their snouts. The ornithopod family, which includes genera such as Edmontosaurus and Parasaurolophus, was a common group of herbivores during the Late Cretaceous Period. Hadrosaurids are descendants of the Upper Jurassic/Lower Cretaceous iguanodontian dinosaurs and had a similar body layout. Hadrosaurs were among the most dominant herbivores during the Late Cretaceous in Asia and North America, and during the close of the Cretaceous several lineages dispersed into Europe, Africa, South America and Antarctica.

Ankylosauria is a group of herbivorous dinosaurs of the order Ornithischia. It includes the great majority of dinosaurs with armor in the form of bony osteoderms, similar to turtles. Ankylosaurs were bulky quadrupeds, with short, powerful limbs. They are known to have first appeared in the Middle Jurassic, and persisted until the end of the Cretaceous Period. The two main families of Ankylosaurs, Nodosauridae and Ankylosauridae are primarily known from the Northern Hemisphere, but the more basal Parankylosauria are known from southern Gondwana during the Cretaceous.

Ceratopsia or Ceratopia is a group of herbivorous, beaked dinosaurs that thrived in what are now North America, Europe, and Asia, during the Cretaceous Period, although ancestral forms lived earlier, in the Jurassic. The earliest known ceratopsian, Yinlong downsi, lived between 161.2 and 155.7 million years ago. The last ceratopsian species, Triceratops prorsus, became extinct during the Cretaceous–Paleogene extinction event, 66 million years ago.

Dromaeosauridae is a family of feathered theropod dinosaurs. They were generally small to medium-sized feathered carnivores that flourished in the Cretaceous Period. The name Dromaeosauridae means 'running lizards', from Greek δρομαῖος (dromaîos), meaning 'running at full speed, swift', and σαῦρος (saûros), meaning 'lizard'. In informal usage, they are often called raptors, a term popularized by the film Jurassic Park; several genera include the term "raptor" directly in their name, and popular culture has come to emphasize their bird-like appearance and speculated bird-like behavior.

Gondwana was a large landmass, often referred to as a supercontinent, that formed during the late Neoproterozoic and began to break up during the Jurassic period. The final stages of break-up, involving the separation of Antarctica from South America and Australia, occurred during the Paleogene. Gondwana was not considered a supercontinent by the earliest definition, since the landmasses of Baltica, Laurentia, and Siberia were separated from it. To differentiate it from the Indian region of the same name, it is also commonly called Gondwanaland.

Abelisauridae is a family of ceratosaurian theropod dinosaurs. Abelisaurids thrived during the Cretaceous period, on the ancient southern supercontinent of Gondwana, and today their fossil remains are found on the modern continents of Africa and South America, as well as on the Indian subcontinent and the island of Madagascar. Isolated teeth were found in the Late Jurassic of Portugal, and the Late Cretaceous genera Tarascosaurus and Arcovenator have been described in France. Abelisaurids first appear in the fossil record of the early middle Jurassic period, and at least three genera survived until the end of the Mesozoic era 66 million years ago.

Carcharodontosauridae is a group of carnivorous theropod dinosaurs. In 1931, Ernst Stromer named Carcharodontosauridae as a family, which, in modern paleontology, indicates a clade within Carnosauria. Carcharodontosaurids include some of the largest land predators ever known: Giganotosaurus, Mapusaurus, Carcharodontosaurus, and Tyrannotitan all rivaled or exceeded Tyrannosaurus in size. A 2015 paper by Christophe Hendrickx and colleagues gives a maximum length estimate of 14 metres (46 ft) for the largest carcharodontosaurids, while the smallest carcharodontosaurids were estimated to have been at least 6 metres (20 ft) long.

The Enantiornithes, also known as enantiornithines or enantiornitheans in literature, are a group of extinct avialans, the most abundant and diverse group known from the Mesozoic era. Almost all retained teeth and clawed fingers on each wing, but otherwise looked much like modern birds externally. Over eighty species of Enantiornithes have been named, but some names represent only single bones, so it is likely that not all are valid. The Enantiornithes became extinct at the Cretaceous–Paleogene boundary, along with Hesperornithes and all other non-avian dinosaurs.

Ornithurae is a natural group which includes the common ancestor of Ichthyornis, Hesperornis, and all modern birds as well as all other descendants of that common ancestor.

Avisaurus is a genus of enantiornithine bird from the Late Cretaceous of North America.

Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae, commonly called angiosperms. The term "angiosperm" is derived from the Greek words ἀγγεῖον /angeion and σπέρμα / sperma ('seed'), meaning those plants that produce their seeds enclosed within a fruit. They are by far the most diverse group of land plants with 64 orders, 416 families, approximately 13,000 known genera and 300,000 known species. Angiosperms were formerly called Magnoliophyta.

Magnolia is a large genus of about 210 to 340 flowering plant species in the subfamily Magnolioideae of the family Magnoliaceae. It is named after French botanist Pierre Magnol.

Sassafras is a genus of three extant and one extinct species of deciduous trees in the family Lauraceae, native to eastern North America and eastern Asia. The genus is distinguished by its aromatic properties, which have made the tree useful to humans.

A rose is either a woody perennial flowering plant of the genus Rosa, in the family Rosaceae, or the flower it bears. There are over three hundred species and tens of thousands of cultivars. They form a group of plants that can be erect shrubs, climbing, or trailing, with stems that are often armed with sharp prickles. Their flowers vary in size and shape and are usually large and showy, in colours ranging from white through yellows and reds. Most species are native to Asia, with smaller numbers native to Europe, North America, and northwestern Africa. Species, cultivars and hybrids are all widely grown for their beauty and often are fragrant. Roses have acquired cultural significance in many societies. Rose plants range in size from compact, miniature roses, to climbers that can reach seven meters in height. Different species hybridize easily, and this has been used in the development of the wide range of garden roses.

Willows, also called sallows and osiers, of the genus Salix, comprise around 350 species of typically deciduous trees and shrubs, found primarily on moist soils in cold and temperate regions.

The Cretaceous–Paleogene (K–Pg) extinction event, also known as the Cretaceous–Tertiary (K–T) extinction, was a sudden mass extinction of three-quarters of the plant and animal species on Earth, approximately 66 million years ago. The event caused the extinction of all non-avian dinosaurs. Most other tetrapods weighing more than 25 kilograms also became extinct, with the exception of some ectothermic species such as sea turtles and crocodilians. It marked the end of the Cretaceous Period, and with it the Mesozoic era, while heralding the beginning of the Cenozoic era, which continues to this day.

An extinction event is a widespread and rapid decrease in the biodiversity on Earth. Such an event is identified by a sharp change in the diversity and abundance of multicellular organisms. It occurs when the rate of extinction increases with respect to the background extinction rate and the rate of speciation. Estimates of the number of major mass extinctions in the last 540 million years range from as few as five to more than twenty. These differences stem from disagreement as to what constitutes a "major" extinction event, and the data chosen to measure past diversity.

The Cretaceous–Paleogene (K–Pg) boundary, formerly known as the Cretaceous–Tertiary (K–T) boundary, is a geological signature, usually a thin band of rock containing much more iridium than other bands. The K–Pg boundary marks the end of the Cretaceous Period, the last period of the Mesozoic Era, and marks the beginning of the Paleogene Period, the first period of the Cenozoic Era. Its age is usually estimated at around 66 million years, with radiometric dating yielding a more precise age of 66.043 ± 0.011 Ma.

The Cenozoic is Earth's current geological era, representing the last 66 million years of Earth's history. It is characterised by the dominance of mammals, birds and flowering plants, a cooling and drying climate, and the current configuration of continents. It is the latest of three geological eras since complex life evolved, preceded by the Mesozoic and Paleozoic. It started with the Cretaceous–Paleogene extinction event, when many species, including the non-avian dinosaurs, became extinct in an event attributed by most experts to the impact of a large asteroid or other celestial body, the Chicxulub impactor.

Birds are a group of warm-blooded vertebrates constituting the class Aves, characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweight skeleton. Birds live worldwide and range in size from the 5.5 cm (2.2 in) bee hummingbird to the 2.8 m common ostrich. There are about ten thousand living species, more than half of which are passerine, or "perching" birds. Birds have wings whose development varies according to species; the only known groups without wings are the extinct moa and elephant birds. Wings, which are modified forelimbs, gave birds the ability to fly, although further evolution has led to the loss of flight in some birds, including ratites, penguins, and diverse endemic island species. The digestive and respiratory systems of birds are also uniquely adapted for flight. Some bird species of aquatic environments, particularly seabirds and some waterbirds, have further evolved for swimming.

Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 245 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is a subject of active research. They became the dominant terrestrial vertebrates after the Triassic–Jurassic extinction event 201.3 mya and their dominance continued throughout the Jurassic and Cretaceous periods. The fossil record shows that birds are feathered dinosaurs, having evolved from earlier theropods during the Late Jurassic epoch, and are the only dinosaur lineage known to have survived the Cretaceous–Paleogene extinction event approximately 66 mya. Dinosaurs can therefore be divided into avian dinosaurs—birds—and the extinct non-avian dinosaurs, which are all dinosaurs other than birds.

Mosasaurs comprise a group of extinct, large marine reptiles from the Late Cretaceous. Their first fossil remains were discovered in a limestone quarry at Maastricht on the Meuse in 1764. They belong to the order Squamata, which includes lizards and snakes.

Invertebrates are a paraphyletic group of animals that neither possess nor develop a vertebral column, derived from the notochord. This is a grouping including all animals apart from the chordate subphylum Vertebrata. Familiar examples of invertebrates include arthropods, mollusks, annelids, echinoderms and cnidarians.

In biological phylogenetics, a clade, also known as a monophyletic group or natural group, is a grouping of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. In the taxonomical literature, sometimes the Latin form cladus is used rather than the English form.