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Jurassic

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Jurassic
201.4 ± 0.2 – ~145.0 Ma
Chronology
Etymology
Name formalityFormal
Usage information
Regional usageGlobal (ICS)
Time scale(s) usedICS Time Scale
Definition
Chronological unitPeriod
Stratigraphic unitSystem
Time span formalityFormal
Lower boundary definitionFirst appearance of the ammonite Psiloceras spelae tirolicum.
Lower boundary GSSPKuhjoch section, Karwendel mountains, Northern Calcareous Alps, Austria
47°29′02″N 11°31′50″E / 47.4839°N 11.5306°E / 47.4839; 11.5306
Lower GSSP ratified2010
Upper boundary definitionNot formally defined
Upper boundary definition candidates
Upper boundary GSSP candidate section(s)None

The Jurassic (/ʊˈræsɪk/ juu-RASS-ik[2]) is a geologic period and stratigraphic system that spanned from the end of the Triassic Period 201.4 million years ago (Mya) to the beginning of the Cretaceous Period, approximately 145 Mya. The Jurassic constitutes the middle period of the Mesozoic Era and is named after the Jura Mountains, where limestone strata from the period were first identified.

The start of the Jurassic was marked by the major Triassic–Jurassic extinction event, associated with the eruption of the Central Atlantic Magmatic Province. The beginning of the Toarcian Stage started around 183 million years ago and is marked by the Toarcian Oceanic Anoxic Event, a global episode of oceanic anoxia, ocean acidification, and elevated temperatures associated with extinctions, likely caused by the eruption of the Karoo-Ferrar large igneous provinces. The end of the Jurassic, however, has no clear boundary with the Cretaceous and is the only boundary between geological periods to remain formally undefined.

By the beginning of the Jurassic, the supercontinent Pangaea had begun rifting into two landmasses: Laurasia to the north and Gondwana to the south. The climate of the Jurassic was warmer than the present, and there were no ice caps. Forests grew close to the poles, with large arid expanses in the lower latitudes.

On land, the fauna transitioned from the Triassic fauna, dominated jointly by dinosauromorph and pseudosuchian archosaurs, to one dominated by dinosaurs alone. The first birds appeared during the Jurassic, evolving from a branch of theropod dinosaurs. Other major events include the appearance of the earliest lizards and the evolution of therian mammals. Crocodylomorphs made the transition from a terrestrial to an aquatic life. The oceans were inhabited by marine reptiles such as ichthyosaurs and plesiosaurs, while pterosaurs were the dominant flying vertebrates. The first sharks, rays and crabs also first appeared during the period.

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Cretaceous

Cretaceous

The Cretaceous is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of the entire Phanerozoic. The name is derived from the Latin creta, "chalk", which is abundant in the latter half of the period. It is usually abbreviated K, for its German translation Kreide.

Jura Mountains

Jura Mountains

The Jura Mountains are a sub-alpine mountain range a short distance north of the Western Alps and mainly demarcate a long part of the French–Swiss border. While the Jura range proper is located in France and Switzerland, the range continues as the Table Jura northeastwards through northern Switzerland and Germany.

Central Atlantic magmatic province

Central Atlantic magmatic province

The Central Atlantic magmatic province (CAMP) is the Earth's largest continental large igneous province, covering an area of roughly 11 million km2. It is composed mainly of basalt that formed before Pangaea broke up in the Mesozoic Era, near the end of the Triassic and the beginning of the Jurassic periods. The subsequent breakup of Pangaea created the Atlantic Ocean, but the massive igneous upwelling provided a legacy of basaltic dikes, sills, and lavas now spread over a vast area around the present central North Atlantic Ocean, including large deposits in northwest Africa, southwest Europe, as well as northeast South America and southeast North America. The name and CAMP acronym were proposed by Andrea Marzoli and adopted at a symposium held at the 1999 Spring Meeting of the American Geophysical Union.

Anoxic event

Anoxic event

Oceanic anoxic events or anoxic events (anoxia conditions) describe periods wherein large expanses of Earth's oceans were depleted of dissolved oxygen (O2), creating toxic, euxinic (anoxic and sulfidic) waters. Although anoxic events have not happened for millions of years, the geologic record shows that they happened many times in the past. Anoxic events coincided with several mass extinctions and may have contributed to them. These mass extinctions include some that geobiologists use as time markers in biostratigraphic dating. On the other hand, there are widespread, various black-shale beds from the mid-Cretaceous which indicate anoxic events but are not associated with mass extinctions. Many geologists believe oceanic anoxic events are strongly linked to the slowing of ocean circulation, climatic warming, and elevated levels of greenhouse gases. Researchers have proposed enhanced volcanism (the release of CO2) as the "central external trigger for euxinia."

Karoo-Ferrar

Karoo-Ferrar

The Karoo and Ferrar Large Igneous Provinces (LIPs) are two large igneous provinces in Southern Africa and Antarctica respectively, collectively known as the Karoo-Ferrar, Gondwana, or Southeast African LIP, associated with the initial break-up of the Gondwana supercontinent at c. 183 Ma. Its flood basalt mostly covers South Africa and Antarctica but portions extend further into southern Africa and into South America, India, Australia and New Zealand.

Laurasia

Laurasia

Laurasia was the more northern of two large landmasses that formed part of the Pangaea supercontinent from around 335 to 175 million years ago (Mya), the other being Gondwana. It separated from Gondwana 215 to 175 Mya during the breakup of Pangaea, drifting farther north after the split and finally broke apart with the opening of the North Atlantic Ocean c. 56 Mya. The name is a portmanteau of Laurentia and Asia.

Gondwana

Gondwana

Gondwana was a large landmass, often referred to as a supercontinent, that formed during the late Neoproterozoic and began to break up during the Jurassic period. The final stages of break-up, involving the separation of Antarctica from South America and Australia, occurred during the Paleogene. Gondwana was not considered a supercontinent by the earliest definition, since the landmasses of Baltica, Laurentia, and Siberia were separated from it. To differentiate it from the Indian region of the same name, it is also commonly called Gondwanaland.

Dinosauromorpha

Dinosauromorpha

Dinosauromorpha is a clade of avemetatarsalian archosaurs that includes the Dinosauria (dinosaurs) and some of their close relatives. It was originally defined to include dinosauriforms and lagerpetids, with later formulations specifically excluding pterosaurs from the group. Birds are the only dinosauromorphs which survive to the present day.

Archosaur

Archosaur

Archosauria is a clade of diapsids, with birds and crocodilians as the only living representatives. Archosaurs are broadly classified as reptiles, in the cladistic sense of the term, which includes birds. Extinct archosaurs include non-avian dinosaurs, pterosaurs, and extinct relatives of crocodilians. Modern paleontologists define Archosauria as a crown group that includes the most recent common ancestor of living birds and crocodilians, and all of its descendants. The base of Archosauria splits into two clades: Pseudosuchia, which includes crocodilians and their extinct relatives, and Avemetatarsalia, which includes birds and their extinct relatives.

Dinosaur

Dinosaur

Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 245 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is a subject of active research. They became the dominant terrestrial vertebrates after the Triassic–Jurassic extinction event 201.3 mya and their dominance continued throughout the Jurassic and Cretaceous periods. The fossil record shows that birds are feathered dinosaurs, having evolved from earlier theropods during the Late Jurassic epoch, and are the only dinosaur lineage known to have survived the Cretaceous–Paleogene extinction event approximately 66 mya. Dinosaurs can therefore be divided into avian dinosaurs—birds—and the extinct non-avian dinosaurs, which are all dinosaurs other than birds.

Crocodylomorpha

Crocodylomorpha

Crocodylomorpha is a group of pseudosuchian archosaurs that includes the crocodilians and their extinct relatives. They were the only members of Pseudosuchia to survive the end-Triassic extinction.

Ichthyosaur

Ichthyosaur

Ichthyosaurs are large extinct marine reptiles. Ichthyosaurs belong to the order known as Ichthyosauria or Ichthyopterygia.

Etymology and history

Portrait of Alexandre Brongniart, who coined the term "Jurassic"
Portrait of Alexandre Brongniart, who coined the term "Jurassic"

The chronostratigraphic term "Jurassic" is linked to the Jura Mountains, a forested mountain range that mainly follows the France–Switzerland border. The name "Jura" is derived from the Celtic root *jor via Gaulish *iuris "wooded mountain", which was borrowed into Latin as a name of a place and evolved into Juria and finally Jura.

During a tour of the region in 1795, German naturalist Alexander von Humboldt recognized carbonate deposits within the Jura Mountains as geologically distinct from the Triassic aged Muschelkalk of Southern Germany, but he erroneously concluded that they were older. He then named them Jura-Kalkstein ('Jura limestone') in 1799.[3]

In 1829, the French naturalist Alexandre Brongniart published a book entitled Description of the Terrains that Constitute the Crust of the Earth or Essay on the Structure of the Known Lands of the Earth. In this book, Brongniart used the phrase terrains jurassiques when correlating the "Jura-Kalkstein" of Humboldt with similarly aged oolitic limestones in Britain, thus coining and publishing the term "Jurassic".[4][3]

The German geologist Leopold von Buch in 1839 established the three-fold division of the Jurassic, originally named from oldest to the youngest: the Black Jurassic, Brown Jurassic, and White Jurassic.[5] The term "Lias" had previously been used for strata of equivalent age to the Black Jurassic in England by William Conybeare and William Phillips in 1822.

The French palaeontologist Alcide d'Orbigny in papers between 1842 and 1852 divided the Jurassic into ten stages based on ammonite and other fossil assemblages in England and France, of which seven are still used, but none has retained its original definition. The German geologist and palaeontologist Friedrich August von Quenstedt in 1858 divided the three series of von Buch in the Swabian Jura into six subdivisions defined by ammonites and other fossils.

The German palaeontologist Albert Oppel in his studies between 1856 and 1858 altered d'Orbigny's original scheme and further subdivided the stages into biostratigraphic zones, based primarily on ammonites. Most of the modern stages of the Jurassic were formalized at the Colloque du Jurassique à Luxembourg in 1962.[3]

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Alexandre Brongniart

Alexandre Brongniart

Alexandre Brongniart was a French chemist, mineralogist, geologist, paleontologist, and zoologist, who collaborated with Georges Cuvier on a study of the geology of the region around Paris. Observing fossil content as well as lithology in sequences, he classified Tertiary formations and was responsible for defining 19th century geological studies as a subject of science by assembling observations and classifications.

Chronostratigraphy

Chronostratigraphy

Chronostratigraphy is the branch of stratigraphy that studies the ages of rock strata in relation to time.

Jura Mountains

Jura Mountains

The Jura Mountains are a sub-alpine mountain range a short distance north of the Western Alps and mainly demarcate a long part of the French–Swiss border. While the Jura range proper is located in France and Switzerland, the range continues as the Table Jura northeastwards through northern Switzerland and Germany.

France–Switzerland border

France–Switzerland border

The France–Switzerland border is 572 km (355 mi) long. Its current path is mostly the product of the Congress of Vienna of 1815, with the accession of Geneva, Neuchâtel and Valais to the Swiss Confederation, but it has since been modified in detail, the last time being in 2002. Although most of the border, marked with border stones, is unguarded, several checkpoints remain staffed, most notably on busy roads.

Celtic languages

Celtic languages

The Celtic languages are a group of related languages descended from Proto-Celtic. They form a branch of the Indo-European language family. The term "Celtic" was first used to describe this language group by Edward Lhuyd in 1707, following Paul-Yves Pezron, who made the explicit link between the Celts described by classical writers and the Welsh and Breton languages.

Gaulish

Gaulish

Gaulish is an extinct Celtic language spoken in parts of Continental Europe before and during the period of the Roman Empire. In the narrow sense, Gaulish was the language of the Celts of Gaul. In a wider sense, it also comprises varieties of Celtic that were spoken across much of central Europe ("Noric"), parts of the Balkans, and Anatolia ("Galatian"), which are thought to have been closely related. The more divergent Lepontic of Northern Italy has also sometimes been subsumed under Gaulish.

Latin

Latin

Latin is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area around present-day Rome, but through the power of the Roman Republic it became the dominant language in the Italian region and subsequently throughout the Roman Empire. Even after the fall of Western Rome, Latin remained the common language of international communication, science, scholarship and academia in Europe until well into the 18th century, when other regional vernaculars supplanted it in common academic and political usage. For most of the time it was used, it would be considered a "dead language" in the modern linguistic definition; that is, it lacked native speakers, despite being used extensively and actively.

Alexander von Humboldt

Alexander von Humboldt

Friedrich Wilhelm Heinrich Alexander von Humboldt was a German polymath, geographer, naturalist, explorer, and proponent of Romantic philosophy and science. He was the younger brother of the Prussian minister, philosopher, and linguist Wilhelm von Humboldt (1767–1835). Humboldt's quantitative work on botanical geography laid the foundation for the field of biogeography. Humboldt's advocacy of long-term systematic geophysical measurement laid the foundation for modern geomagnetic and meteorological monitoring.

Carbonate rock

Carbonate rock

Carbonate rocks are a class of sedimentary rocks composed primarily of carbonate minerals. The two major types are limestone, which is composed of calcite or aragonite (different crystal forms of CaCO3), and dolomite rock (also known as dolostone), which is composed of mineral dolomite (CaMg(CO3)2).

Christian Leopold von Buch

Christian Leopold von Buch

Christian Leopold von Buch, usually cited as Leopold von Buch, was a German geologist and paleontologist born in Stolpe an der Oder and is remembered as one of the most important contributors to geology in the first half of the nineteenth century. His scientific interest was devoted to a broad spectrum of geological topics: volcanism, petrology, fossils, stratigraphy and mountain formation. His most remembered accomplishment is the scientific definition of the Jurassic system.

Black Jurassic

Black Jurassic

The Black Jurassic or Black Jura in earth history refers to the lowest of the three lithostratigraphic units of the South German Jurassic, the latter being understood not as a geographical, but a geological term in the sense of a lithostratigraphic super group. Formerly and even occasionally today in the popular scientific literature, this term is equated to the chronostratigraphic series of the Lower Jurassic. This is however not quite correct, because the Black Jura does not exactly coincide with the chronostratigraphic boundaries of the Lower Jurassic. In addition, the term lias, which was formerly frequently used as a synonym, should no longer be used in connexion with the South German Jurassic. Instead, the term lias, or North German Lias, should be reserved for the roughly equivalent lithostratigraphic unit in the North German Jurassic. The deposition of the Black Jurassic took place about 199 to 175 million years ago. The Black Jurassic follows the lithostratigraphic unit of the Keuper and is superimposed on the lithostratigraphic group of the Brown Jurassic.

Brown Jurassic

Brown Jurassic

The Brown Jurassic or Brown Jura in earth history refers to the middle of the three lithostratigraphic units of the South German Jurassic, the latter being understood not as a geographical, but a geological term in the sense of a lithostratigraphic super group. Formerly and even occasionally today in the popular scientific literature, this term is equated to the chronostratigraphic series of the Middle Jurassic. This is however not quite correct, because the Brown Jura does not exactly coincide with the chronostratigraphic boundaries of the Middle Jurassic. In addition, the term Dogger, which was formerly frequently used as a synonym should no longer be used in connexion with the South German Jurassic. Instead the term "Dogger" should be reserved for the roughly equivalent lithostratigraphic unit in the North German Jurassic. The Brown Jurassic was deposited about 175-161 million years ago. The Brown Jurassic follows the lithostratigraphic unit of the Black Jurassic and is superimposed on the lithostratigraphic group of the White Jurassic.

Geology

The Jurassic Period is divided into three epochs: Early, Middle, and Late. Similarly, in stratigraphy, the Jurassic is divided into the Lower Jurassic, Middle Jurassic, and Upper Jurassic series. Geologists divide the rocks of the Jurassic into a stratigraphic set of units called stages, each formed during corresponding time intervals called ages.

Stages can be defined globally or regionally. For global stratigraphic correlation, the International Commission on Stratigraphy (ICS) ratify global stages based on a Global Boundary Stratotype Section and Point (GSSP) from a single formation (a stratotype) identifying the lower boundary of the stage.[3] The ages of the Jurassic from youngest to oldest are as follows:[6]

Series/epoch Stage/age Lower boundary
Early Cretaceous Berriasian ~145 Mya
Upper/Late Jurassic Tithonian 152.1 ±0.9 Mya
Kimmeridgian 157.3 ±1.0 Mya
Oxfordian 163.5 ±1.0 Mya
Middle Jurassic Callovian 166.1 ±1.2 Mya
Bathonian 168.3 ±1.3 Mya
Bajocian 170.3 ±1.4 Mya
Aalenian 174.1 ±1.0 Mya
Lower/Early Jurassic Toarcian 182.7 ±0.7 Mya
Pliensbachian 190.8 ±1.0 Mya
Sinemurian 199.3 ±0.3 Mya
Hettangian 201.3 ±0.2 Mya

Stratigraphy

Folded Lower Jurassic limestone layers of the Doldenhorn nappe at Gasteretal, Switzerland
Folded Lower Jurassic limestone layers of the Doldenhorn nappe at Gasteretal, Switzerland
Middle Jurassic strata in Neuquén Province, Argentina
Middle Jurassic strata in Neuquén Province, Argentina
Tidwell Member of the Morrison Formation (Upper Jurassic), Colorado
Tidwell Member of the Morrison Formation (Upper Jurassic), Colorado

Jurassic stratigraphy is primarily based on the use of ammonites as index fossils. The first appearance datum of specific ammonite taxa is used to mark the beginnings of stages, as well as smaller timespans within stages, referred to as "ammonite zones"; these, in turn, are also sometimes subdivided further into subzones. Global stratigraphy is based on standard European ammonite zones, with other regions being calibrated to the European successions.[3]

Base Aalenian GSSP at Fuentelsaz
Base Aalenian GSSP at Fuentelsaz

Early Jurassic

The oldest part of the Jurassic Period has historically been referred to as the Lias or Liassic, roughly equivalent in extent to the Early Jurassic, but also including part of the preceding Rhaetian. The Hettangian Stage was named by Swiss palaeontologist Eugène Renevier in 1864 after Hettange-Grande in north-eastern France.[3] The GSSP for the base of the Hettangian is located at the Kuhjoch Pass, Karwendel Mountains, Northern Calcareous Alps, Austria; it was ratified in 2010. The beginning of the Hettangian, and thus the Jurassic as a whole, is marked by the first appearance of the ammonite Psiloceras spelae tirolicum in the Kendlbach Formation exposed at Kuhjoch.[7] The base of the Jurassic was previously defined as the first appearance of Psiloceras planorbis by Albert Oppel in 1856–58, but this was changed as the appearance was seen as too localised an event for an international boundary.[3]

The Sinemurian Stage was first defined and introduced into scientific literature by Alcide d'Orbigny in 1842. It takes its name from the French town of Semur-en-Auxois, near Dijon. The original definition of Sinemurian included what is now the Hettangian. The GSSP of the Sinemurian is located at a cliff face north of the hamlet of East Quantoxhead, 6 kilometres east of Watchet, Somerset, England, within the Blue Lias, and was ratified in 2000. The beginning of the Sinemurian is defined by the first appearance of the ammonite Vermiceras quantoxense.[3][8]

Albert Oppel in 1858 named the Pliensbachian Stage after the hamlet of Pliensbach in the community of Zell unter Aichelberg in the Swabian Alb, near Stuttgart, Germany. The GSSP for the base of the Pliensbachian is found at the Wine Haven locality in Robin Hood's Bay, Yorkshire, England, in the Redcar Mudstone Formation, and was ratified in 2005. The beginning of the Pliensbachian is defined by the first appearance of the ammonite Bifericeras donovani.[9]

The village Thouars (Latin: Toarcium), just south of Saumur in the Loire Valley of France, lends its name to the Toarcian Stage. The Toarcian was named by Alcide d'Orbigny in 1842, with the original locality being Vrines quarry around 2 km northwest of Thouars. The GSSP for the base of the Toarcian is located at Peniche, Portugal, and was ratified in 2014. The boundary is defined by the first appearance of ammonites belonging to the subgenus Dactylioceras (Eodactylites).[10]

Middle Jurassic

The Aalenian is named after the city of Aalen in Germany. The Aalenian was defined by Swiss geologist Karl Mayer-Eymar in 1864. The lower boundary was originally between the dark clays of the Black Jurassic and the overlying clayey sandstone and ferruginous oolite of the Brown Jurassic sequences of southwestern Germany.[3] The GSSP for the base of the Aalenian is located at Fuentelsaz in the Iberian range near Guadalajara, Spain, and was ratified in 2000. The base of the Aalenian is defined by the first appearance of the ammonite Leioceras opalinum.[11]

Alcide d'Orbigny in 1842 named the Bajocian Stage after the town of Bayeux (Latin: Bajoce) in Normandy, France. The GSSP for the base of the Bajocian is located in the Murtinheira section at Cabo Mondego, Portugal; it was ratified in 1997. The base of the Bajocian is defined by the first appearance of the ammonite Hyperlioceras mundum.[12]

The Bathonian is named after the city of Bath, England, introduced by Belgian geologist d'Omalius d'Halloy in 1843, after an incomplete section of oolitic limestones in several quarries in the region. The GSSP for the base of the Bathonian is Ravin du Bès, Bas-Auran area, Alpes de Haute Provence, France; it was ratified in 2009. The base of the Bathonian is defined by the first appearance of the ammonite Gonolkites convergens, at the base of the Zigzagiceras zigzag ammonite zone.[13]

The Callovian is derived from the Latinized name of the village of Kellaways in Wiltshire, England, and was named by Alcide d'Orbigny in 1852, originally the base at the contact between the Forest Marble Formation and the Cornbrash Formation. However, this boundary was later found to be within the upper part of the Bathonian.[3] The base of the Callovian does not yet have a certified GSSP. The working definition for the base of the Callovian is the first appearance of ammonites belonging to the genus Kepplerites.[14]

Upper Jurassic

The Oxfordian is named after the city of Oxford in England and was named by Alcide d'Orbigny in 1844 in reference to the Oxford Clay. The base of the Oxfordian lacks a defined GSSP. W. J. Arkell in studies in 1939 and 1946 placed the lower boundary of the Oxfordian as the first appearance of the ammonite Quenstedtoceras mariae (then placed in the genus Vertumniceras). Subsequent proposals have suggested the first appearance of Cardioceras redcliffense as the lower boundary.[3][14]

The village of Kimmeridge on the coast of Dorset, England, is the origin of the name of the Kimmeridgian. The stage was named by Alcide d'Orbigny in 1842 in reference to the Kimmeridge Clay. The GSSP for the base of the Kimmeridgian is the Flodigarry section at Staffin Bay on the Isle of Skye, Scotland,[15] which was ratified in 2021. The boundary is defined by the first appearance of ammonites marking the boreal Bauhini Zone and the subboreal Baylei Zone.[14]

The Tithonian was introduced in scientific literature by Albert Oppel in 1865. The name Tithonian is unusual in geological stage names because it is derived from Greek mythology rather than a place name. Tithonus was the son of Laomedon of Troy and fell in love with Eos, the Greek goddess of dawn. His name was chosen by Albert Oppel for this stratigraphical stage because the Tithonian finds itself hand in hand with the dawn of the Cretaceous. The base of the Tithonian currently lacks a GSSP.[3] The working definition for the base of the Tithonian is the first appearance of the ammonite genus Gravesia.[14]

The upper boundary of the Jurassic is currently undefined, and the Jurassic–Cretaceous boundary is currently the only system boundary to lack a defined GSSP. Placing a GSSP for this boundary has been difficult because of the strong regionality of most biostratigraphic markers, and lack of any chemostratigraphic events, such as isotope excursions (large sudden changes in ratios of isotopes), that could be used to define or correlate a boundary. Calpionellids, an enigmatic group of planktonic protists with urn-shaped calcitic tests briefly abundant during the latest Jurassic to earliest Cretaceous, have been suggested to represent the most promising candidates for fixing the Jurassic–Cretaceous boundary[16] In particular, the first appearance Calpionella alpina, co-inciding with the base of the eponymous Alpina subzone, has been proposed as the definition of the base of the Cretaceous.[17] The working definition for the boundary has often been placed as the first appearance of the ammonite Strambergella jacobi, formerly placed in the genus Berriasella, but its use as a stratigraphic indicator has been questioned, as its first appearance does not correlate with that of C. alpina.[18]

Mineral and hydrocarbon deposits

The Kimmeridge Clay and equivalents are the major source rock for the North Sea oil.[19] The Arabian Intrashelf Basin, deposited during the Middle and Late Jurassic, is the setting of the world's largest oil reserves, including the Ghawar Field, the world's largest oil field.[20] The Jurassic-aged Sargelu[21] and Naokelekan formations[22] are major source rocks for oil in Iraq. Over 1500 gigatons of Jurassic coal reserves are found in north-west China, primarily in the Turpan-Hami Basin and the Ordos Basin.[23]

Impact structures

Major impact structures include the Morokweng impact structure, a 70 km diameter impact structure buried beneath the Kalahari desert in northern South Africa. The impact is dated to the Tithonian, approximately 146.06 ± 0.16 Mya.[24] Another major structure is the Puchezh-Katunki crater, 40 kilometres in diameter, buried beneath Nizhny Novgorod Oblast in western Russia. The impact has been dated to the Sinemurian, 195.9 ± 1.0 Ma.[25]

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Stratigraphy

Stratigraphy

Stratigraphy is a branch of geology concerned with the study of rock layers (strata) and layering (stratification). It is primarily used in the study of sedimentary and layered volcanic rocks. Stratigraphy has three related subfields: lithostratigraphy, biostratigraphy, and chronostratigraphy.

Middle Jurassic

Middle Jurassic

The Middle Jurassic is the second epoch of the Jurassic Period. It lasted from about 174.1 to 163.5 million years ago. Fossils of land-dwelling animals, such as dinosaurs, from the Middle Jurassic are relatively rare, but geological formations containing land animal fossils include the Forest Marble Formation in England, the Kilmaluag Formation in Scotland, the Daohugou Beds in China, the Itat Formation in Russia, and the Isalo III Formation of western Madagascar.

Series (stratigraphy)

Series (stratigraphy)

Series are subdivisions of rock layers based on the age of the rock and formally defined by international conventions of the geological timescale. A series is therefore a sequence of strata defining a chronostratigraphic unit. Series are subdivisions of systems and are themselves divided into stages.

Stage (stratigraphy)

Stage (stratigraphy)

In chronostratigraphy, a stage is a succession of rock strata laid down in a single age on the geologic timescale, which usually represents millions of years of deposition. A given stage of rock and the corresponding age of time will by convention have the same name, and the same boundaries.

International Commission on Stratigraphy

International Commission on Stratigraphy

The International Commission on Stratigraphy (ICS), sometimes unofficially referred to as the "International Stratigraphic Commission", is a daughter or major subcommittee grade scientific daughter organization that concerns itself with stratigraphical, geological, and geochronological matters on a global scale.

Global Boundary Stratotype Section and Point

Global Boundary Stratotype Section and Point

A Global Boundary Stratotype Section and Point (GSSP) is an internationally agreed upon reference point on a stratigraphic section which defines the lower boundary of a stage on the geologic time scale. The effort to define GSSPs is conducted by the International Commission on Stratigraphy, a part of the International Union of Geological Sciences. Most, but not all, GSSPs are based on paleontological changes. Hence GSSPs are usually described in terms of transitions between different faunal stages, though far more faunal stages have been described than GSSPs. The GSSP definition effort commenced in 1977. As of 2022, 78 of the 101 stages that need a GSSP have a ratified GSSP.

Stratotype

Stratotype

A stratotype or type section in geology is the physical location or outcrop of a particular reference exposure of a stratigraphic sequence or stratigraphic boundary. If the stratigraphic unit is layered, it is called a stratotype, whereas the standard of reference for unlayered rocks is the type locality.

Early Cretaceous

Early Cretaceous

The Early Cretaceous or the Lower Cretaceous, is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 145 Ma to 100.5 Ma.

Berriasian

Berriasian

In the geological timescale, the Berriasian is an age/stage of the Early/Lower Cretaceous. It is the oldest subdivision in the entire Cretaceous. It has been taken to span the time between 145.0 ± 4.0 Ma and 139.8 ± 3.0 Ma. The Berriasian succeeds the Tithonian and precedes the Valanginian.

Late Jurassic

Late Jurassic

The Late Jurassic is the third epoch of the Jurassic Period, and it spans the geologic time from 163.5 ± 1.0 to 145.0 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic strata.

Kimmeridgian

Kimmeridgian

In the geologic timescale, the Kimmeridgian is an age in the Late Jurassic Epoch and a stage in the Upper Jurassic Series. It spans the time between 157.3 ± 1.0 Ma and 152.1 ± 0.9 Ma. The Kimmeridgian follows the Oxfordian and precedes the Tithonian.

Callovian

Callovian

In the geologic timescale, the Callovian is an age and stage in the Middle Jurassic, lasting between 165.3 ± 1.1 Ma and 161.5 ± 1.0 Ma. It is the last stage of the Middle Jurassic, following the Bathonian and preceding the Oxfordian.

Paleogeography and tectonics

Pangaea at the start of Jurassic
Pangaea at the start of Jurassic
The breakup of Gondwanaland took place during the Late Jurassic, the Indian Ocean opened up as a result
The breakup of Gondwanaland took place during the Late Jurassic, the Indian Ocean opened up as a result

At the beginning of the Jurassic, all of the world's major landmasses were coalesced into the supercontinent Pangaea, which during the Early Jurassic began to break up into northern supercontinent Laurasia and the southern supercontinent Gondwana.[26] The rifting between North America and Africa was the first to initiate, beginning in the early Jurassic, associated with the emplacement of the Central Atlantic Magmatic Province.[27]

During the Jurassic, the North Atlantic Ocean remained relatively narrow, while the South Atlantic did not open until the Cretaceous.[28][27] The continents were surrounded by Panthalassa, with the Tethys Ocean between Gondwana and Asia. At the end of the Triassic, there was a marine transgression in Europe, flooding most parts of central and western Europe transforming it into an archipelago of islands surrounded by shallow seas.[29] During the Jurassic, both the North and South Pole were covered by oceans.[26] Beginning in the Early Jurassic, the Boreal Ocean was connected to the proto-Atlantic by the "Viking corridor" or Transcontinental Laurasian Seaway, a passage between the Baltic Shield and Greenland several hundred kilometers wide.[30][31]

Madagascar and Antarctica began to rift away from Africa during the late Early Jurassic in association with the eruption of the Karoo-Ferrar large igneous provinces, opening the western Indian Ocean and beginning the fragmentation of Gondwana.[32][33] At the beginning of the Jurassic, North and South America remained connected, but by the beginning of the Late Jurassic they had rifted apart to form the Caribbean Seaway, also known as the Hispanic Corridor, which connected the North Atlantic Ocean with eastern Panthalassa. Palaeontological data suggest that the seaway had been open since the Early Jurassic.[34]

As part of the Nevadan orogeny, which began during the Triassic, the Cache Creek Ocean closed, and various terranes including the large Wrangellia Terrane accreted onto the western margin of North America.[35][36] By the Middle Jurassic the Siberian plate and the North China-Amuria block had collided, resulting in the closure of the Mongol-Okhotsk Ocean.[37]

Formation of the Pacific Plate during the Early Jurassic
Formation of the Pacific Plate during the Early Jurassic

During the Early Jurassic, around 190 million years ago, the Pacific Plate originated at the triple junction of the Farallon, Phoenix, and Izanagi tectonic plates, the three main oceanic plates of Panthalassa. The previously stable triple junction had converted to an unstable arrangement surrounded on all sides by transform faults because of a kink in one of the plate boundaries, resulting in the formation of the Pacific Plate at the centre of the junction.[38] During the Middle to early Late Jurassic, the Sundance Seaway, a shallow epicontinental sea, covered much of northwest North America.[39]

Grainstone with calcitic ooids and sparry calcite cement; Carmel Formation, Middle Jurassic, of southern Utah, USA
Grainstone with calcitic ooids and sparry calcite cement; Carmel Formation, Middle Jurassic, of southern Utah, USA

The eustatic sea level is estimated to have been close to present levels during the Hettangian and Sinemurian, rising several tens of metres during the late Sinemurian–Pliensbachian before regressing to near present levels by the late Pliensbachian. There seems to have been a gradual rise to a peak of ~75 m above present sea level during the Toarcian. During the latest part of the Toarcian, the sea level again dropped by several tens of metres. It progressively rose from the Aalenian onwards, aside from dips of a few tens of metres in the Bajocian and around the Callovian–Oxfordian boundary, peaking possibly as high as 140 metres above present sea level at the Kimmeridgian–Tithonian boundary. The sea levels falls in the late Tithonian, perhaps to around 100 metres, before rebounding to around 110 metres at the Tithonian–Berriasian boundary.

The sea level within the long-term trends across the Jurassic was cyclical, with 64 fluctuations, 15 of which were over 75 metres. The most noted cyclicity in Jurassic rocks is fourth order, with a periodicity of approximately 410,000 years.[40]

During the Early Jurassic the world's oceans transitioned from an aragonite sea to a calcite sea chemistry, favouring the dissolution of aragonite and precipitation of calcite.[41] The rise of calcareous plankton during the Middle Jurassic profoundly altered ocean chemistry, with the deposition of biomineralized plankton on the ocean floor acting as a buffer against large CO2 emissions.[42]

Discover more about Paleogeography and tectonics related topics

Indian Ocean

Indian Ocean

The Indian Ocean is the third-largest of the world's five oceanic divisions, covering 70,560,000 km2 (27,240,000 sq mi) or ~19.8% of the water on Earth's surface. It is bounded by Asia to the north, Africa to the west and Australia to the east. To the south it is bounded by the Southern Ocean or Antarctica, depending on the definition in use. Along its core, the Indian Ocean has some large marginal or regional seas such as the Arabian Sea, Laccadive Sea, Bay of Bengal, and Andaman Sea.

Laurasia

Laurasia

Laurasia was the more northern of two large landmasses that formed part of the Pangaea supercontinent from around 335 to 175 million years ago (Mya), the other being Gondwana. It separated from Gondwana 215 to 175 Mya during the breakup of Pangaea, drifting farther north after the split and finally broke apart with the opening of the North Atlantic Ocean c. 56 Mya. The name is a portmanteau of Laurentia and Asia.

Gondwana

Gondwana

Gondwana was a large landmass, often referred to as a supercontinent, that formed during the late Neoproterozoic and began to break up during the Jurassic period. The final stages of break-up, involving the separation of Antarctica from South America and Australia, occurred during the Paleogene. Gondwana was not considered a supercontinent by the earliest definition, since the landmasses of Baltica, Laurentia, and Siberia were separated from it. To differentiate it from the Indian region of the same name, it is also commonly called Gondwanaland.

Central Atlantic magmatic province

Central Atlantic magmatic province

The Central Atlantic magmatic province (CAMP) is the Earth's largest continental large igneous province, covering an area of roughly 11 million km2. It is composed mainly of basalt that formed before Pangaea broke up in the Mesozoic Era, near the end of the Triassic and the beginning of the Jurassic periods. The subsequent breakup of Pangaea created the Atlantic Ocean, but the massive igneous upwelling provided a legacy of basaltic dikes, sills, and lavas now spread over a vast area around the present central North Atlantic Ocean, including large deposits in northwest Africa, southwest Europe, as well as northeast South America and southeast North America. The name and CAMP acronym were proposed by Andrea Marzoli and adopted at a symposium held at the 1999 Spring Meeting of the American Geophysical Union.

Atlantic Ocean

Atlantic Ocean

The Atlantic Ocean is the second-largest of the world's five oceans, with an area of about 106,460,000 km2 (41,100,000 sq mi). It covers approximately 20% of Earth's surface and about 29% of its water surface area. It is known to separate the "Old World" of Africa, Europe, and Asia from the "New World" of the Americas in the European perception of the World.

Marine transgression

Marine transgression

A marine transgression is a geologic event during which sea level rises relative to the land and the shoreline moves toward higher ground, which results in flooding. Transgressions can be caused by the land sinking or by the ocean basins filling with water or decreasing in capacity. Transgressions and regressions may be caused by tectonic events such as orogenies, severe climate change such as ice ages or isostatic adjustments following removal of ice or sediment load.

Baltic Shield

Baltic Shield

The Baltic Shield is a segment of the Earth's crust belonging to the East European Craton, representing a large part of Fennoscandia, northwestern Russia and the northern Baltic Sea. It is composed mostly of Archean and Proterozoic gneisses and greenstone which have undergone numerous deformations through tectonic activity. It contains the oldest rocks of the European continent with a thickness of 250–300 km.

Greenland

Greenland

Greenland is an island country in North America and part of the Kingdom of Denmark. It lies between the Arctic and Atlantic oceans, east of the Canadian Arctic Archipelago. Greenland is the world's largest island. It is one of three countries that form the Kingdom of Denmark, the others being Denmark and the Faroe Islands; the citizens of all these countries are citizens of Denmark and of the European Union. The capital of Greenland is Nuuk.

Karoo-Ferrar

Karoo-Ferrar

The Karoo and Ferrar Large Igneous Provinces (LIPs) are two large igneous provinces in Southern Africa and Antarctica respectively, collectively known as the Karoo-Ferrar, Gondwana, or Southeast African LIP, associated with the initial break-up of the Gondwana supercontinent at c. 183 Ma. Its flood basalt mostly covers South Africa and Antarctica but portions extend further into southern Africa and into South America, India, Australia and New Zealand.

Nevadan orogeny

Nevadan orogeny

The Nevadan orogeny occurred along the western margin of North America during the Middle Jurassic to Early Cretaceous time which is approximately from 155 Ma to 145 Ma. Throughout the duration of this orogeny there were at least two different kinds of orogenic processes occurring. During the early stages of orogenesis an "Andean type" continental magmatic arc developed due to subduction of the Farallon oceanic plate beneath the North American Plate. The latter stages of orogenesis, in contrast, saw multiple oceanic arc terranes accreted onto the western margin of North America in a "Cordilleran type" accretionary orogen. Deformation related to the accretion of these volcanic arc terranes is mostly limited to the western regions of the resulting mountain ranges and is absent from the eastern regions. In addition, the deformation experienced in these mountain ranges is mostly due to the Nevadan orogeny and not other external events such as the more recent Sevier and Laramide Orogenies. It is noted that the Klamath Mountains and the Sierra Nevada share similar stratigraphy indicating that they were both formed by the Nevadan orogeny. In comparison with other orogenic events, it appears that the Nevadan Orogeny occurred rather quickly taking only about 10 million years as compared to hundreds of millions of years for other orogenies around the world.

Cache Creek Ocean

Cache Creek Ocean

The Cache Creek Ocean, formerly called Anvil Ocean, is an inferred ancient ocean which existed between western North America and offshore continental terranes between the Devonian and the Middle Jurassic.

Accretion (geology)

Accretion (geology)

Accretion, in geology, is a process by which material is added to a tectonic plate at a subduction zone, frequently on the edge of existing continental landmasses. The added material may be sediment, volcanic arcs, seamounts, oceanic crust or other igneous features.

Climate

The climate of the Jurassic was generally warmer than that of present, by around 5 °C to 10 °C, with atmospheric carbon dioxide likely four times higher. Intermittent "cold snap" intervals are known to have occurred during this time period, however, interrupting the otherwise warm greenhouse climate.[43] Forests likely grew near the poles, where they experienced warm summers and cold, sometimes snowy winters; there were unlikely to have been ice sheets given the high summer temperatures that prevented the accumulation of snow, though there may have been mountain glaciers.[44] Dropstones and glendonites in northeastern Siberia during the Early to Middle Jurassic indicate cold winters.[45] The ocean depths were likely 8 °C warmer than present, and coral reefs grew 10° of latitude further north and south. The Intertropical Convergence Zone likely existed over the oceans, resulting in large areas of desert and scrubland in the lower latitudes between 40° N and S of the equator. Tropical rainforest and tundra biomes are likely to have been rare or absent.[44]

The beginning of the Jurassic was likely marked by a thermal spike corresponding to the Triassic–Jurassic extinction and eruption of the Central Atlantic magmatic province. The first part of the Jurassic was marked by the Early Jurassic Cool Interval between 199 and 183 million years ago.[45] It has been proposed that glaciation was present in the Northern Hemisphere during the late Pliensbachian.[46][47] There was a spike in global temperatures of around 4–8 °C during the early part of the Toarcian corresponding to the Toarcian Oceanic Anoxic Event and the eruption of the Karoo-Ferrar large igneous provinces in southern Gondwana, with the warm interval extending to the end of the Toarcian around 174 million years ago.[45]

During the Toarcian Warm Interval, ocean surface temperatures likely exceeded 30 °C, and equatorial and subtropical (30°N–30°S) regions are likely to have been extremely arid, with temperatures in the interior of Pangea likely in excess of 40 °C. The Toarcian Warm Interval is followed by the Middle Jurassic Cool Interval between 174 and 164 million years ago,[45] with the Callovian-Oxfordian boundary at the end of this interval witnessing particularly notable global cooling,[48] potentially even an ice age.[49] This is followed by the Kimmeridgian Warm Interval between 164 and 150 million years ago. The Pangaean interior had less severe seasonal swings than in previous warm periods as the expansion of the Central Atlantic and Western Indian Ocean provided new sources of moisture.[45] A prominent drop in temperatures occurred during the Tithonian, known as the Early Tithonian Cooling Event (ETCE).[48] The end of the Jurassic was marked by the Tithonian–early Barremian Cool Interval, beginning 150 million years ago and continuing into the Early Cretaceous.[45]

Climatic events

Toarcian Oceanic Anoxic Event

The Toarcian Oceanic Anoxic Event (TOAE), also known as the Jenkyns Event, was an episode of widespread oceanic anoxia during the early part of the Toarcian Age, c. 183 Mya. It is marked by a globally documented high amplitude negative carbon isotope excursion,[50][51] as well as the deposition of black shales[52] and the extinction and collapse of carbonate-producing marine organisms, associated with a major rise in global temperatures.[53]

The TOAE is often attributed to the eruption of the Karoo-Ferrar large igneous provinces and the associated increase of carbon dioxide concentration in the atmosphere, as well as the possible associated release of methane clathrates.[53] This likely accelerated the hydrological cycle and increased silicate weathering, as evidenced by an increased amount of organic matter of terrestrial origin found in marine deposits during the TOAE.[54] Groups affected include ammonites,[55] ostracods,[52][56] foraminifera,[57][58] bivalves,[52] cnidarians, and especially brachiopods,[59][60][61] for which the TOAE represented one of the most severe extinctions in their evolutionary history.[62] While the event had significant impact on marine invertebrates, it had little effect on marine reptiles.[63] During the TOAE, the Sichuan Basin was transformed into a giant lake, probably three times the size of modern-day Lake Superior, represented by the Da’anzhai Member of the Ziliujing Formation. The lake likely sequestered ∼460 gigatons (Gt) of organic carbon and ∼1,200 Gt of inorganic carbon during the event.[64] Seawater pH, which had already substantially decreased prior to the event, increased slightly during the early stages of the TOAE, before dropping to its lowest point around the middle of the event.[65] This ocean acidification is the probable cause of the collapse of carbonate production.[66][67] Additionally, anoxic conditions were exacerbated by enhanced recycling of phosphorus back into ocean water as a result of high ocean acidity and temperature inhibiting its mineralisation into apatite; the abundance of phosphorus in marine environments caused further eutrophication and consequent anoxia in a positive feedback loop.[68]

End-Jurassic transition

The end-Jurassic transition was originally considered one of eight mass extinctions, but is now considered to be a complex interval of faunal turnover, with the increase in diversity of some groups and decline in others, though the evidence for this is primarily European, probably controlled by changes in eustatic sea level.[69]

Discover more about Climate related topics

Carbon dioxide in Earth's atmosphere

Carbon dioxide in Earth's atmosphere

In Earth's atmosphere, carbon dioxide is a trace gas that plays an integral part in the greenhouse effect, carbon cycle, photosynthesis and oceanic carbon cycle. It is one of several greenhouse gases in the atmosphere of Earth. The current global average concentration of CO2 in the atmosphere is 421 ppm as of May 2022. This is an increase of 50% since the start of the Industrial Revolution, up from 280 ppm during the 10,000 years prior to the mid-18th century. The increase is due to human activity. Burning fossil fuels is the main cause of these increased CO2 concentrations and also the main cause of climate change. Other large anthropogenic sources include cement production, deforestation, and biomass burning.

Dropstone

Dropstone

Dropstones are isolated fragments of rock found within finer-grained water-deposited sedimentary rocks or pyroclastic beds. They range in size from small pebbles to boulders. The critical distinguishing feature is that there is evidence that they were not transported by normal water currents, but rather dropped in vertically through the air or water column. Such deposition can occur i.e. during a volcanic eruption.

Coral reef

Coral reef

A coral reef is an underwater ecosystem characterized by reef-building corals. Reefs are formed of colonies of coral polyps held together by calcium carbonate. Most coral reefs are built from stony corals, whose polyps cluster in groups.

Intertropical Convergence Zone

Intertropical Convergence Zone

The Intertropical Convergence Zone, known by sailors as the doldrums or the calms because of its monotonous windless weather, is the area where the northeast and the southeast trade winds converge. It encircles Earth near the thermal equator though its specific position varies seasonally. When it lies near the geographic Equator, it is called the near-equatorial trough. Where the ITCZ is drawn into and merges with a monsoonal circulation, it is sometimes referred to as a monsoon trough, a usage that is more common in Australia and parts of Asia.

Large igneous province

Large igneous province

A large igneous province (LIP) is an extremely large accumulation of igneous rocks, including intrusive and extrusive, arising when magma travels through the crust towards the surface. The formation of LIPs is variously attributed to mantle plumes or to processes associated with divergent plate tectonics. The formation of some of the LIPs in the past 500 million years coincide in time with mass extinctions and rapid climatic changes, which has led to numerous hypotheses about causal relationships. LIPs are fundamentally different from any other currently active volcanoes or volcanic systems.

Anoxic event

Anoxic event

Oceanic anoxic events or anoxic events (anoxia conditions) describe periods wherein large expanses of Earth's oceans were depleted of dissolved oxygen (O2), creating toxic, euxinic (anoxic and sulfidic) waters. Although anoxic events have not happened for millions of years, the geologic record shows that they happened many times in the past. Anoxic events coincided with several mass extinctions and may have contributed to them. These mass extinctions include some that geobiologists use as time markers in biostratigraphic dating. On the other hand, there are widespread, various black-shale beds from the mid-Cretaceous which indicate anoxic events but are not associated with mass extinctions. Many geologists believe oceanic anoxic events are strongly linked to the slowing of ocean circulation, climatic warming, and elevated levels of greenhouse gases. Researchers have proposed enhanced volcanism (the release of CO2) as the "central external trigger for euxinia."

Methane clathrate

Methane clathrate

Methane clathrate (CH4·5.75H2O) or (8CH4·46H2O), also called methane hydrate, hydromethane, methane ice, fire ice, natural gas hydrate, or gas hydrate, is a solid clathrate compound (more specifically, a clathrate hydrate) in which a large amount of methane is trapped within a crystal structure of water, forming a solid similar to ice. Originally thought to occur only in the outer regions of the Solar System, where temperatures are low and water ice is common, significant deposits of methane clathrate have been found under sediments on the ocean floors of the Earth. Methane hydrate is formed when hydrogen-bonded water and methane gas come into contact at high pressures and low temperatures in oceans.

Carbonate–silicate cycle

Carbonate–silicate cycle

The carbonate–silicate geochemical cycle, also known as the inorganic carbon cycle, describes the long-term transformation of silicate rocks to carbonate rocks by weathering and sedimentation, and the transformation of carbonate rocks back into silicate rocks by metamorphism and volcanism. Carbon dioxide is removed from the atmosphere during burial of weathered minerals and returned to the atmosphere through volcanism. On million-year time scales, the carbonate-silicate cycle is a key factor in controlling Earth's climate because it regulates carbon dioxide levels and therefore global temperature.

Ostracod

Ostracod

Ostracods, or ostracodes, are a class of the Crustacea, sometimes known as seed shrimp. Some 70,000 species have been identified, grouped into several orders. They are small crustaceans, typically around 1 mm (0.039 in) in size, but varying from 0.2 to 30 mm in the case of Gigantocypris. Their bodies are flattened from side to side and protected by a bivalve-like, chitinous or calcareous valve or "shell". The hinge of the two valves is in the upper (dorsal) region of the body. Ostracods are grouped together based on gross morphology. While early work indicated the group may not be monophyletic and early molecular phylogeny was ambiguous on this front, recent combined analyses of molecular and morphological data found support for monophyly in analyses with broadest taxon sampling.

Foraminifera

Foraminifera

Foraminifera are single-celled organisms, members of a phylum or class of amoeboid protists characterized by streaming granular ectoplasm for catching food and other uses; and commonly an external shell of diverse forms and materials. Tests of chitin are believed to be the most primitive type. Most foraminifera are marine, the majority of which live on or within the seafloor sediment, while a smaller number float in the water column at various depths, which belong to the suborder Globigerinina. Fewer are known from freshwater or brackish conditions, and some very few (nonaquatic) soil species have been identified through molecular analysis of small subunit ribosomal DNA.

Bivalvia

Bivalvia

Bivalvia, in previous centuries referred to as the Lamellibranchiata and Pelecypoda, is a class of marine and freshwater molluscs that have laterally compressed bodies enclosed by a shell consisting of two hinged parts. As a group, bivalves have no head and they lack some usual molluscan organs, like the radula and the odontophore. The class includes the clams, oysters, cockles, mussels, scallops, and numerous other families that live in saltwater, as well as a number of families that live in freshwater. The majority are filter feeders. The gills have evolved into ctenidia, specialised organs for feeding and breathing. Most bivalves bury themselves in sediment, where they are relatively safe from predation. Others lie on the sea floor or attach themselves to rocks or other hard surfaces. Some bivalves, such as the scallops and file shells, can swim. The shipworms bore into wood, clay, or stone and live inside these substances.

Cnidaria

Cnidaria

Cnidaria is a phylum under kingdom Animalia containing over 11,000 species of aquatic animals found both in freshwater and marine environments, predominantly the latter.

Flora

End-Triassic extinction

There is no evidence of a mass extinction of plants at the Triassic–Jurassic boundary.[70] At the Triassic–Jurassic boundary in Greenland, the sporomorph (pollen and spores) record suggests a complete floral turnover.[71] An analysis of macrofossil floral communities in Europe suggests that changes were mainly due to local ecological succession.[72] At the end of the Triassic, the Peltaspermaceae became extinct in most parts of the world, with Lepidopteris persisting into the Early Jurassic in Patagonia.[73] Dicroidium, a seed fern that was a dominant part of Gondwanan floral communities during the Triassic, also declined at the Triassic–Jurassic boundary, surviving as a relict in Antarctica into the Sinemurian.[74]

Floral composition

Conifers

Petrified Araucaria mirabilis cone from the Middle Jurassic of Argentina
Petrified Araucaria mirabilis cone from the Middle Jurassic of Argentina

Conifers formed a dominant component of Jurassic floras. The Late Triassic and Jurassic was a major time of diversification of conifers, with most modern conifer groups appearing in the fossil record by the end of the Jurassic, having evolved from voltzialean ancestors.[75][76]

Araucarian conifers have their first unambiguous records during the Early Jurassic, and members of the modern genus Araucaria were widespread across both hemispheres by the Middle Jurassic.[76][77][78]

Also abundant during the Jurassic is the extinct family Cheirolepidiaceae, often recognised through their highly distinctive Classopolis pollen. Jurassic representatives include the pollen cone Classostrobus and the seed cone Pararaucaria. Araucarian and Cheirolepidiaceae conifers often occur in association.[79]

The oldest definitive record of the cypress family (Cupressaceae) is Austrohamia minuta from the Early Jurassic (Pliensbachian) of Patagonia, known from many parts of the plant.[80] The reproductive structures of Austrohamia have strong similarities to those of the primitive living cypress genera Taiwania and Cunninghamia. By the Middle to Late Jurassic Cupressaceae were abundant in warm temperate–tropical regions of the Northern Hemisphere, most abundantly represented by the genus Elatides.[81]

Members of the extinct genus Schizolepidopsis which likely represent a stem-group to the pine family (Pinaceae), were widely distributed across Eurasia during the Jurassic.[82][83] The oldest unambiguous record of Pinaceae is the pine cone Eathiestrobus, known from the Late Jurassic (Kimmeridgian) of Scotland, which remains the only known unequivocal fossil of the group before the Cretaceous.[84] Despite being the earliest known member of the Pinaceae, Eathiestrobus appears to be a member of the pinoid clade of the family, suggesting that the initial diversification of Pinaceae occurred earlier than has been found in the fossil record.[85][76]

During the Early Jurassic, the flora of the mid-latitudes of Eastern Asia were dominated by the extinct deciduous broad leafed conifer Podozamites, which appears to not be closely related to any living family of conifer. Its range extended northwards into polar latitudes of Siberia and then contracted northward in the Middle to Late Jurassic, corresponding to the increasing aridity of the region.[86]

The earliest record of the yew family (Taxaceae) is Palaeotaxus rediviva, from the Hettangian of Sweden, suggested to be closely related to the living Austrotaxus, while Marskea jurassica from the Middle Jurassic of Yorkshire, England and material from the Callovian–Oxfordian Daohugou Bed in China are thought to be closely related to Amentotaxus, with the latter material assigned to the modern genus, indicating that Taxaceae had substantially diversified by the end of the Jurassic.[87]

Podocarpaceae, today largely confined to the Southern Hemisphere, occurred in the Northern Hemisphere during the Jurassic, Examples include Podocarpophyllum from the Early to Middle Jurassic of Central Asia and Siberia,[88] Scarburgia from the Middle Jurassic of Yorkshire,[89] and Harrisiocarpus from the Jurassic of Poland.[90]

Ginkgoales

Leaves of Ginkgo huttonii from the Middle Jurassic of England
Leaves of Ginkgo huttonii from the Middle Jurassic of England

Ginkgoales, of which the sole living species is Ginkgo biloba, were more diverse during the Jurassic: they were among the most important components of Eurasian Jurassic floras and were adapted to a wide variety of climatic conditions.[91] The earliest representatives of the genus Ginkgo, represented by ovulate and pollen organs similar to those of the modern species, are known from the Middle Jurassic in the Northern Hemisphere.[91] Several other lineages of ginkgoaleans are known from Jurassic rocks, including Yimaia, Grenana, Nagrenia and Karkenia. These lineages are associated with Ginkgo-like leaves, but are distinguished from living and fossil representatives of Ginkgo by having differently arranged reproductive structures.[91][92] Umaltolepis from the Jurassic of Asia has strap-shaped ginkgo-like leaves with highly distinct reproductive structures with similarities to those of peltasperm and corystosperm seed ferns, has been suggested to be a member of Ginkgoales sensu lato.[93]

Bennettitales

Restoration of a member of Bennettitales belonging to Williamsoniaceae.
Restoration of a member of Bennettitales belonging to Williamsoniaceae.

Bennettitales, having first become widespread during the preceding Triassic, were diverse and abundant members of Jurassic floras across both hemispheres.[94] The foliage of Bennettitales bears strong similarities to those of cycads, to such a degree that they cannot be reliably distinguished on the basis of morphology alone. Leaves of Bennettitales can be distinguished from those of cycads their different arrangement of stomata, and the two groups are not thought to be closely related.[95] Jurassic Bennettitales predominantly belong to the group Williamsoniaceae,[94] which grew as shrubs and small trees. The Williamsoniaceae are thought to have had a divaricate branching habit, similar to that of living Banksia, and adapted to growing in open habitats with poor soil nutrient conditions.[96] Bennettitales exhibit complex, flower-like reproductive structures some of which are thought to have been pollinated by insects. Several groups of insects that bear long proboscis, including extinct families such as kalligrammatid lacewings[97] and extant ones such as acrocerid flies,[98] are suggested to have been pollinators of bennettitales, feeding on nectar produced by bennettitalean cones.

Cycads

Cycads reached their apex of diversity during the Jurassic and Cretaceous Periods.[99] Despite the Mesozoic sometimes being called the "Age of Cycads", cycads are thought to have been a relatively minor component of mid-Mesozoic floras, with the Bennettitales and Nilssoniales, which have cycad-like foliage, being dominant.[100] The Nilssoniales have often been considered cycads or cycad relatives, but have been found to be distinct on chemical grounds, and perhaps more closely allied with Bennettitales.[101] Cycads are thought to have been mostly confined to tropical and subtropical latitudes throughout their evolutionary history.[101] The relationships of most Mesozoic cycads to living groups are ambiguous.[100] Modern cycads are pollinated by beetles, and such an association is thought to have formed by the Early Jurassic.[99]

Other seed plants

Although there have been several claimed records and phylogenetic stem group age estimates for individual early diverging angiosperm orders, there are no widely accepted Jurassic fossil records of flowering plants, which make up 90% of living plant species, and fossil evidence suggests that the group diversified during the following Cretaceous.[102]

The earliest known Gnetophytes appeared by the end of the Jurassic.[103][104]

Sagenopteris phillipsi (Caytoniales) from the Middle Jurassic of Yorkshire, England
Sagenopteris phillipsi (Caytoniales) from the Middle Jurassic of Yorkshire, England

"Seed ferns" (Pteridospermatophyta) is a collective term to refer to disparate lineages of fern like plants that produce seeds but have uncertain affinities to living seed plant groups. A prominent group of Jurassic seed ferns is the Caytoniales, which reached their zenith during the Jurassic, with widespread records in the Northern Hemisphere, though records in the Southern Hemisphere remain rare. Due to their berry-like seed-bearing capsules, they have often been suggested to have been closely related or perhaps ancestral to flowering plants, but the evidence for this is inconclusive.[105] A variety of other Jurassic seed ferns of uncertain placement are known, including Pachypteris from Europe, which has sometimes been allied with the corystosperms.[106]

Czekanowskiales, also known as Leptostrobales, are a group of seed plants uncertain affinities with persistent heavily dissected leaves borne on deciduous short shoots, subtended by scale-like leaves, known from the Late Triassic (possibly Late Permian[107]) to Cretaceous.[108] They are thought to have had a tree- or shrub-like habit and formed a conspicuous component of Northern Hemisphere Mesozoic temperate and warm-temperate floras.[107] The genus Phoenicopsis was widespread in Early-Middle Jurassic floras of Eastern Asia and Siberia.[109]

The Pentoxylales, a small but clearly distinct group of liana-like seed plants of obscure affinities, first appeared during the Jurassic. Their distribution appears to have been confined to Eastern Gondwana.[110]

Ferns and allies

Living families of ferns widespread during the Jurassic include Dipteridaceae, Matoniaceae, Gleicheniaceae, Osmundaceae and Marattiaceae.[111][112] Polypodiales, which make up 80% of living fern diversity, have no record from the Jurassic and are thought to have diversified in the Cretaceous,[113] though the widespread Jurassic herbaceous fern genus Coniopteris, historically interpreted as a close relative of tree ferns of the family Dicksoniaceae, has recently been reinterpreted as an early relative of the group.[114]

The Cyatheales, the group containing most modern tree ferns, appeared during the Late Jurassic, represented by members of the genus Cyathocaulis, which are suggested to be early members of Cyatheaceae on the basis of cladistic analysis.[115] Only a handful of possible records exist of the Hymenophyllaceae from the Jurassic, including Hymenophyllites macrosporangiatus from the Russian Jurassic.[116]

The oldest remains of modern horsetails of the genus Equisetum first appear in the Early Jurassic, represented by Equisetum dimorphum from the Early Jurassic of Patagonia[117] and Equisetum laterale from the Early to Middle Jurassic of Australia.[118][119] Silicified remains of Equisetum thermale from the Late Jurassic of Argentina exhibit all the morphological characters of modern members of the genus.[120] The estimated split between Equisetum bogotense and all other living Equisetum is estimated to have occurred no later than the Early Jurassic.[119]

Lower plants

Quillworts virtually identical to modern species are known from the Jurassic onwards. Isoetites rolandii from the Middle Jurassic of Oregon is the earliest known species to represent all major morphological features of modern Isoetes. More primitive forms such as Nathorstiana, which retain an elongated stem, persisted into the Early Cretaceous.[121]

The moss Kulindobryum from the Middle Jurassic of Russia, which was found associated with dinosaur bones, is thought to be related to the Splachnaceae, which grow on animal caracasses.[122] Bryokhutuliinia from the same region is thought to be related to Dicranales.[122] Heinrichsiella from the Jurassic of Patagonia is thought to belong to either Polytrichaceae or Timmiellaceae.[123]

The liverwort Pellites hamiensis from the Middle Jurassic Xishanyao Formation of China is the oldest record of the family Pelliaceae.[124] Pallaviciniites sandaolingensis from the same deposit is thought to belong to the subclass Pallaviciniineae within the Pallaviciniales.[125] Ricciopsis sandaolingensis, also from the same deposit, is the only Jurassic record of Ricciaceae.[126]

Discover more about Flora related topics

Ecological succession

Ecological succession

Ecological succession is the process of change in the species structure of an ecological community over time. The time scale can be decades or more or less.

Dicroidium

Dicroidium

Dicroidium is an extinct genus of fork-leaved seed plants assigned to the order Umkomasiales (corystosperms) that were widely distributed over Gondwana during the Triassic. Their fossils are known from South Africa, the Arabian Peninsula, Australia, New Zealand, South America, Madagascar, the Indian subcontinent and Antarctica. They were first discovered in Triassic sediments of Tasmania by Morris in 1845. Fossils from the Umm Irna Formation in Jordan and in Pakistan indicate that these plants already existed in Late Permian. Late surviving members of the genus are known from the Early Jurassic (Sinemurian) of East Antarctica. Within paleobotany, Dicroidium is a form genus used to refers to the leaves, associated with ovuluate organs classified as Umkomasia and pollen organs classified as Pteruchus, while Dicroidum is also used collectively to refer to the whole plant.

Araucaria mirabilis

Araucaria mirabilis

Araucaria mirabilis is an extinct species of coniferous tree from Patagonia, Argentina. It belongs to the genus Araucaria.

Conifer

Conifer

Conifers are a group of cone-bearing seed plants, a subset of gymnosperms. Scientifically, they make up the division Pinophyta, also known as Coniferophyta or Coniferae. The division contains a single extant class, Pinopsida. All extant conifers are perennial woody plants with secondary growth. The great majority are trees, though a few are shrubs. Examples include cedars, Douglas-firs, cypresses, firs, junipers, kauri, larches, pines, hemlocks, redwoods, spruces, and yews. As of 1998, the division Pinophyta was estimated to contain eight families, 68 genera, and 629 living species.

Araucariaceae

Araucariaceae

Araucariaceae – also known as araucarians – is a family of coniferous trees. The family achieved its maximum diversity during the Jurassic and Cretaceous periods, when it was distributed almost worldwide. Most of the Araucariaceae in the Northern Hemisphere vanished in the Cretaceous–Paleogene extinction event, and they are now largely confined to the Southern Hemisphere, except for a few species of Agathis in Southeast Asia.

Araucaria

Araucaria

Araucaria is a genus of evergreen coniferous trees in the family Araucariaceae. There are 20 extant species in New Caledonia, Norfolk Island, eastern Australia, New Guinea, East Argentina, South Brazil, Chile and Paraguay. They are still common in the South Pacific region and Eastern Australia.

Cheirolepidiaceae

Cheirolepidiaceae

Cheirolepidiaceae is an extinct family of conifers. They first appeared in the Triassic, and were widespread during most of the Mesozoic era. They are united by the possession of a distinctive pollen type assigned to the form genus Classopollis. The name Frenelopsidaceae or "frenelopsids" has been used for a group of Cheirolepidiaceae with jointed stems, thick internode cuticles, sheathing leaf bases and reduced free leaf tips. The leaf morphology has been noted as being similar to that of halophyte Salicornia. Several members of the family appear to have been adapted for semi-arid and coastal settings, with a high tolerance of saline conditions. Cheirolepidiaceae disappeared from most regions of the world during the Cenomanian-Turonian stages of the Late Cretaceous, but reappeared in South America during the Maastrichtian, the final stage of the Cretaceous, increasing in abundance after the K-Pg extinction and being a prominent part of the regional fauna during the Paleocene, before going extinct.

Cupressaceae

Cupressaceae

Cupressaceae is a conifer family, the cypress family, with worldwide distribution. The family includes 27–30 genera, which include the junipers and redwoods, with about 130–140 species in total. They are monoecious, subdioecious or (rarely) dioecious trees and shrubs up to 116 m (381 ft) tall. The bark of mature trees is commonly orange- to red- brown and of stringy texture, often flaking or peeling in vertical strips, but smooth, scaly or hard and square-cracked in some species.

Cunninghamia

Cunninghamia

Cunninghamia is a genus of one or two living species of evergreen coniferous trees in the cypress family Cupressaceae. They are native to China, northern Vietnam and Laos, and perhaps also Cambodia. They may reach 50 m (160 ft) in height. In vernacular use, it is most often known as Cunninghamia, but is also sometimes called "China-fir". The genus name Cunninghamia honours Dr. James Cunningham, a British doctor who introduced this species into cultivation in 1702 and botanist Allan Cunningham.

Conifer cone

Conifer cone

A conifer cone is a seed-bearing organ on gymnosperm plants. It is usually woody, ovoid to globular, including scales and bracts arranged around a central axis, especially in conifers and cycads. The cone of Pinophyta contains the reproductive structures. The woody cone is the female cone, which produces seeds. The male cone, which produces pollen, is usually herbaceous and much less conspicuous even at full maturity. The name "cone" derives from Greek konos, which also gave name to the geometric cone. The individual plates of a cone are known as scales. The umbo of a conifer cone refers to the first year's growth of a seed scale on the cone, showing up as a protuberance at the end of the two-year-old scale.

Eathiestrobus

Eathiestrobus

Eathiestrobus mackenziei is a fossil pine cone found in the Kimmeridge Clay Formation near Eathie, on the Black Isle in Scotland. It is the oldest fossil pine currently known.

Clade

Clade

In biological phylogenetics, a clade, also known as a monophyletic group or natural group, is a grouping of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. In the taxonomical literature, sometimes the Latin form cladus is used rather than the English form.

Fauna

Reptiles

Crocodylomorphs

The Triassic–Jurassic extinction decimated pseudosuchian diversity, with crocodylomorphs, which originated during the early Late Triassic, being the only group of pseudosuchians to survive, with all others, including the herbivorous aetosaurs and carnivorous "rauisuchians" becoming extinct.[127] The morphological diversity of crocodylomorphs during the Early Jurassic was around the same as those of Late Triassic pseudosuchians, but they occupied different areas of morphospace, suggesting that they occupied different ecological niches to their Triassic counterparts and that there was an extensive and rapid radiation of crocodylomorphs during this interval.[128] While living crocodilians are confined to an aquatic ambush predator lifestyle, Jurassic crocodylomorphs exhibited a wide variety of life habits. An unnamed protosuchid known from teeth from the Early Jurassic of Arizona represents the earliest known herbivorous crocodylomorph, an adaptation that appeared several times during the Mesozoic.[129]

The Thalattosuchia, a clade of predominantly marine crocodylomorphs, first appeared during the Early Jurassic and became a prominent part of marine ecosystems.[130] Within Thalattosuchia, the Metriorhynchidae became highly adapted for life in the open ocean, including the transformation of limbs into flippers, the development of a tail fluke, and smooth, scaleless skin.[131] The morphological diversity of crocodylomorphs during the Early and Middle Jurassic was relatively low compared to that in later time periods and was dominated by terrestrial small-bodied, long-legged sphenosuchians, early crocodyliforms and thalattosuchians.[132][130] The Neosuchia, a major group of crocodylomorphs, first appeared during the Early to Middle Jurassic. The Neosuchia represents the transition from an ancestrally terrestrial lifestyle to a freshwater aquatic ecology similar to that occupied by modern crocodilians.[133] The timing of the origin of Neosuchia is disputed. The oldest record of Neosuchians has been suggested to be Calsoyasuchus, from the Early Jurassic of Arizona, which in many analyses has been recovered as the earliest branching member of the neosuchian family Goniopholididae, which radically alters times of diversification for crocodylomorphs. However, this placement has been disputed, with some analyses finding it outside Neosuchia, which would place the oldest records of Neosuchia in the Middle Jurassic.[133] Razanandrongobe from the Middle Jurassic of Madagascar has been suggested the represent the oldest record of Notosuchia, a primarily Gondwanan clade of mostly terrestrial crocodylomorphs, otherwise known from the Cretaceous and Cenozoic.[134]

Turtles

Thalassemys, a thalassochelydian sea turtle known from the Late Jurassic of Germany
Thalassemys, a thalassochelydian sea turtle known from the Late Jurassic of Germany

Stem-group turtles (Testudinata) diversified during the Jurassic. Jurassic stem-turtles belong to two progressively more advanced clades, the Mesochelydia and Perichelydia.[135] It is thought that the ancestral condition for mesochelydians is aquatic, as opposed to terrestrial for testudinates.[136] The two modern groups of turtles (Testudines), Pleurodira and Cryptodira, diverged by the beginning of the Late Jurassic.[135] The oldest known pleurodires, the Platychelyidae, are known from the Late Jurassic of Europe and the Americas,[137] while the oldest unambiguous cryptodire, Sinaspideretes, an early relative of softshell turtles, is known from the Late Jurassic of China.[138] The Thalassochelydia, a diverse lineage of marine turtles unrelated to modern sea turtles, are known from the Late Jurassic of Europe and South America.[139]

Lepidosaurs

Rhynchocephalians (the sole living representative being the tuatara) had achieved a global distribution by the beginning of the Jurassic.[140] Rhynchocephalians reached their highest morphological diversity in their evolutionary history during the Jurassic, occupying a wide range of lifestyles, including the aquatic pleurosaurs with long snake-like bodies and reduced limbs, the specialized herbivorous eilenodontines, as well as Oenosaurus, which had broad tooth plates indicative of durophagy.[141] Rhynchocephalians disappeared from Asia after the Early Jurassic.[140] The last common ancestor of living squamates (which includes lizards and snakes) is estimated to have lived around 190 million years ago during the Early Jurassic, with the major divergences between modern squamate lineages estimated to have occurred during the Early to Middle Jurassic.[142] Squamates first appear in the fossil record during the Middle Jurassic[143] including members of modern clades such as Scincomorpha,[144] though many Jurassic squamates have unclear relationships to living groups.[145] Eichstaettisaurus from the Late Jurassic of Germany has been suggested to be an early relative of geckos and displays adaptations for climbing.[146] Dorsetisaurus from the Late Jurassic of North America and Europe represents the oldest widely accepted record of Anguimorpha.[147] Tamaulipasaurus from Early Jurassic of Mexico and Marmoretta from the Middle Jurassic of Britain represents late surviving lepidosauromorphs outside both Rhynchocephalia and Squamata.[148]

Choristoderes

Skeleton of Coeruleodraco
Skeleton of Coeruleodraco

The earliest known remains of Choristodera, a group of freshwater aquatic reptiles with uncertain affinities to other reptile groups, are found in the Middle Jurassic. Only two genera of choristodere are known from the Jurassic. One is the small lizard-like Cteniogenys, thought to be the most basal known choristodere; it is known from the Middle to Late Jurassic of Europe and Late Jurassic of North America, with similar remains also known from the upper Middle Jurassic of Kyrgyzstan and western Siberia.[149] The other is Coeruleodraco from the Late Jurassic of China, which is a more advanced choristodere, though still small and lizard-like in morphology.[150]

Ichthyosaurs

Ichthyosaurs suffered an evolutionary bottleneck during the end-Triassic extinction, with all non-neoichthyosaurians becoming extinct. Ichthyosaurs reached their apex of species diversity during the Early Jurassic, with an array of morphologies including the huge apex predator Temnodontosaurus and swordfish-like Eurhinosaurus, though Early Jurassic ichthyosaurs were significantly less morphologically diverse than their Triassic counterparts.[151][152] At the Early–Middle Jurassic boundary, between the end of the Toarcian and the beginning of the Bajocian, most lineages of ichythosaur appear to have become extinct, with the first appearance of the Ophthalmosauridae, the clade that would encompass almost all ichthyosaurs from then on, during the early Bajocian.[153] Ophthalmosaurids were diverse by the Late Jurassic, but failed to fill many of the niches that had been occupied by ichthyosaurs during the Early Jurassic.[153][151][152]

Plesiosaurs

Rhomaleosaurus cramptoni at the Natural History Museum
Rhomaleosaurus cramptoni at the Natural History Museum

Plesiosaurs originated at the end of the Triassic (Rhaetian). By the end of the Triassic, all other sauropterygians, including placodonts and nothosaurs, had become extinct. At least six lineages of plesiosaur crossed the Triassic–Jurassic boundary.[154] Plesiosaurs were already diverse in the earliest Jurassic, with the majority of plesiosaurs in the Hettangian-aged Blue Lias belonging to the Rhomaleosauridae. Early plesiosaurs were generally small-bodied, with body size increasing into the Toarcian.[155] There appears to have been a strong turnover around the Early–Middle Jurassic boundary, with microcleidids and rhomaleosaurids becoming extinct and nearly extinct respectively after the end of the Toarcian with the first appearance of the dominant clade of plesiosaurs of the latter half of the Jurassic, the Cryptoclididae during the Bajocian.[153] The Middle Jurassic saw the evolution of short-necked and large-headed thalassophonean pliosaurs from ancestrally small-headed, long-necked forms.[156][153] Some thalassophonean pliosaurs, such as some species of Pliosaurus, had skulls up to two metres in length with body lengths estimated around 10–12 metres, making them the apex predators of Late Jurassic oceans.[157][153] Plesiosaurs invaded freshwater environments during the Jurassic, with indeterminate remains of small-bodied pleisosaurs known from freshwater sediments from the Jurassic of China and Australia.[158][159]

Pterosaurs

Pterosaurs first appeared in the Late Triassic. A major radiation of Jurassic pterosaurs is the Rhamphorhynchidae, which first appeared in the late Early Jurassic (Toarcian);[160] they are thought to been piscivorous.[161] Anurognathids, which first appeared in the Middle Jurassic, possessed short heads and densely furred bodies, and are thought to have been insectivores.[161] Derived monofenestratan pterosaurs such as wukongopterids appeared in the late Middle Jurassic. Advanced short-tailed pterodactyloids first appeared at the Middle–Late Jurassic boundary. Jurassic pterodactyloids include the ctenochasmatids, like Ctenochasma, which have closely spaced needle-like teeth that were presumably used for filter feeding.[161] The bizarre Late Jurassic ctenochasmatoid Cycnorhamphus had a jaw with teeth only at the tips, with bent jaws like those of living openbill storks that may have been used to hold and crush hard invertebrates.[161]

Dinosaurs

Dinosaurs, which had morphologically diversified in the Late Triassic, experienced a major increase in diversity and abundance during the Early Jurassic in the aftermath of the end-Triassic extinction and the extinction of other reptile groups, becoming the dominant vertebrates in terrestrial ecosystems.[162][163] Chilesaurus, a morphologically aberrant herbivorous dinosaur from the Late Jurassic of South America, has uncertain relationships to the three main groups of dinosaurs, having been recovered as a member of all three in different analyses.[164]

Theropods

Advanced theropods belonging to Neotheropoda first appeared in the Late Triassic. Basal neotheropods, such as coelophysoids and dilophosaurs, persisted into the Early Jurassic, but became extinct by the Middle Jurassic.[165] The earliest averostrans appear during the Early Jurassic, with the earliest known member of Ceratosauria being Saltriovenator from the early Sinemurian (199.3–197.5 million years ago) of Italy.[166] The unusual ceratosaur Limusaurus from the Late Jurassic of China had a herbivorous diet, with adults having edentulous beaked jaws,[167] making it the earliest known theropod to have converted from an ancestrally carnivorous diet.[168] The earliest members of the Tetanurae appeared during the late Early Jurassic or early Middle Jurassic.[169] The Megalosauridae represent the oldest radiation of the Tetanurae, first appearing in Europe during the Bajocian.[170] The oldest member of Allosauroidea has been suggested to be Asfaltovenator from the Middle Jurassic of South America.[169] Coelurosaurs first appeared during the Middle Jurassic, including early tyrannosaurs such as Proceratosaurus from the Bathonian of Britain.[171] Some coelurosaurs from the Late Jurassic of China including Shishugounykus and Haplocheirus are suggested to represent early alvarezsaurs,[172] however, this has been questioned.[173] Scansoriopterygids, a group of small feathered coelurosaurs with membraneous, bat-like wings for gliding, are known from the Middle to Late Jurassic of China.[174] The oldest record of troodontids is suggested to be Hesperornithoides from the Late Jurassic of North America. Tooth remains suggested to represent those of dromaeosaurs are known from the Jurassic, but no body remains are known until the Cretaceous.[175]

Birds
Archaeopteryx lithographica from the Late Jurassic (Tithonian) of Germany
Archaeopteryx lithographica from the Late Jurassic (Tithonian) of Germany

The earliest avialans, which include birds and their ancestors, appear during the Middle to Late Jurassic, definitively represented by Archaeopteryx from the Late Jurassic of Germany. Avialans belong to the clade Paraves within Coelurosauria, which also includes dromaeosaurs and troodontids. The Anchiornithidae from the Middle-Late Jurassic of Eurasia have frequently suggested to be avialans, but have also alternatively found as a separate lineage of paravians.[176]

Skeleton of Heterodontosaurus, a primitive ornithischian from the Early Jurassic of South Africa
Skeleton of Heterodontosaurus, a primitive ornithischian from the Early Jurassic of South Africa

Ornithischians

The earliest definitive ornithischians appear during the Early Jurassic, represented by basal ornithischians like Lesothosaurus, heterodontosaurids, and early members of Thyreophora. The earliest members of Ankylosauria and Stegosauria appear during the Middle Jurassic.[177] The basal neornithischian Kulindadromeus from the Middle Jurassic of Russia indicates that at least some ornithischians were covered in protofeathers.[178] The earliest members of Ankylopollexia, which become prominent in the Cretaceous, appeared during the Late Jurassic, represented by bipedal forms such as Camptosaurus.[179] Ceratopsians first appeared in the Late Jurassic of China, represented by members of Chaoyangsauridae.[180]

Sauropodomorphs

Skeleton of Mamenchisaurus sinocanadorum from the Middle-Late Jurassic of China
Skeleton of Mamenchisaurus sinocanadorum from the Middle-Late Jurassic of China

Sauropods became the dominant large herbivores in terrestrial ecosystems during the Jurassic.[181] Some Jurassic sauropods reached gigantic sizes, becoming the largest organisms to have ever lived on land.[182]

Basal bipedal sauropodomorphs, such as massospondylids, continued to exist into the Early Jurassic, but became extinct by the beginning of the Middle Jurassic.[181] Quadrupedal sauropomorphs appeared during the Late Triassic. The quadrupedal Ledumahadi from the earliest Jurassic of South Africa reached an estimated weight of 12 tons, far in excess of other known basal sauropodomorphs.[183] Gravisaurian sauropods first appeared during the Early Jurassic, with the oldest definitive record being Vulcanodon from Zimbabwe, likely of Sinemurian age.[184] Eusauropods first appeared during the late Early Jurassic (Toarcian) and diversified during the Middle Jurassic;[181] these included cetiosaurids, turiasaurs,[185] and mamenchisaurs.[186] Neosauropods such as macronarians and diplodocoids first appeared during the Middle Jurassic, before becoming abundant and globally distributed during the Late Jurassic.[187]

Amphibians

Skeleton of Karaurus sharovi, a stem-group salamander from the Middle to Late Jurassic of Kazakhstan
Skeleton of Karaurus sharovi, a stem-group salamander from the Middle to Late Jurassic of Kazakhstan

The diversity of temnospondyls had progressively declined through the Late Triassic, with only brachyopoids surviving into the Jurassic and beyond.[188] Members of the family Brachyopidae are known from Jurassic deposits in Asia,[189] while the chigutisaurid Siderops is known from the Early Jurassic of Australia.[190] Modern lissamphibians began to diversify during the Jurassic. The Early Jurassic Prosalirus thought to represent the first frog relative with a morphology capable of hopping like living frogs.[191] Morphologically recognisable stem-frogs like the South American Notobatrachus are known from the Middle Jurassic,[192] with modern crown-group frogs like Enneabatrachus and Rhadinosteus appearing by the Late Jurassic.[193] While the earliest salamander-line amphibians are known from the Triassic,[194] crown group salamanders first appear during the Middle to Late Jurassic in Eurasia, alongside stem-group relatives. Many Jurassic stem-group salamanders, such as Marmorerpeton and Kokartus, are thought to have been neotenic.[195] Early representatives of crown group salamanders include Chunerpeton, Pangerpeton and Linglongtriton from the Middle to Late Jurassic Yanliao Biota of China. These belong to the Cryptobranchoidea, which contains living Asiatic and giant salamanders.[196] Beiyanerpeton, and Qinglongtriton from the same biota are thought to be early members of Salamandroidea, the group which contains all other living salamanders.[197][198] Salamanders dispersed into North America by the end of the Jurassic, as evidenced by Iridotriton, found in the Late Jurassic Morrison Formation.[199] The oldest undisputed stem-caecilian is the Early Jurassic Eocaecilia from Arizona.[200] The fourth group of lissamphibians, the extinct albanerpetontids, first appeared in the Middle Jurassic, represented by Anoualerpeton priscus from the Bathonian of Britain, as well as indeterminate remains from equivalently aged sediments in France and the Anoual Formation of Morocco.[201]

Henkelotherium, a likely arboreal dyolestoid from the Late Jurassic of Portugal
Henkelotherium, a likely arboreal dyolestoid from the Late Jurassic of Portugal

Mammaliaformes

Mammaliaformes, having originated from cynodonts at the end of the Triassic, diversified extensively during the Jurassic. Important groups of Jurassic Mammaliaformes include Morganucodonta, Docodonta, Eutriconodonta, Dryolestida, Haramiyida and Multituberculata. While most Jurassic mammalaliaformes are solely known from isolated teeth and jaw fragments, exceptionally preserved remains have revealed a variety of lifestyles.[202] The docodontan Castorocauda was adapted to aquatic life, similarly to the platypus and otters.[203] Some members of Haramiyida[204] and the eutriconodontan tribe Volaticotherini[205] had a patagium akin to those of flying squirrels, allowing them to glide through the air. The aardvark-like mammal Fruitafossor, of uncertain taxonomy, was likely a specialist on colonial insects, similarly to living anteaters.[206] Australosphenida, a group of mammals possibly related to monotremes, first appeared in the Middle Jurassic of Gondwana.[207] Therian mammals, represented today by living placentals and marsupials, appear during the early Late Jurassic, represented by Juramaia, a eutherian mammal closer to the ancestry of placentals than marsupials.[208] Juramaia is much more advanced than expected for its age, as other therian mammals are not known until the Early Cretaceous.[209] Two groups of non-mammalian cynodonts persisted beyond the end of the Triassic. The insectiviorous Tritheledontidae has a few records from the Early Jurassic. The Tritylodontidae, a herbiviorous group of cynodonts that first appeared during the Rhaetian, has abundant records from the Jurassic, overwhelmingly from the Northern Hemisphere.[210][211]

Fish

Conodonts

The last known species of conodont, a class of jawless fish whose hard, tooth-like elements are key index fossils, finally became extinct during the earliest Jurassic after over 300 million years of evolutionary history, with an asynchronous extinction occurring first in the Tethys and eastern Panthalassa and survivors persisting into the earliest Hettangian of Hungary and central Panthalassa.[212] End-Triassic conodonts were represented by only a handful of species and had been progressively declining through the Middle and Late Triassic.[213]

Sarcopterygii

Coelacanth from the Solnhofen Limestone
Coelacanth from the Solnhofen Limestone

Lungfish (Dipnoi) were present in freshwater environments of both hemispheres during the Jurassic. Genera include Ceratodus and Ptychoceratodus, which are more closely related to living South American and African lungfish than Queensland lungfish, and Ferganoceratodus from the Jurassic of Asia, which is not closely related to either group of living lungfish.[214] Mawsoniids, a marine and freshwater/brackish group of coelacanths, which first appeared in North America during the Triassic, expanded into Europe and South America by the end of the Jurassic.[215] The marine Latimeriidae, which contains the living coelacanths of the genus Latimeria, were also present in the Jurassic, having originated in the Triassic.[216]

Actinopterygii

Fossil of Thrissops, an ichthyodectid stem-group teleost from the Late Jurassic Solnhofen Limestone of Germany, showing preserved colouration
Fossil of Thrissops, an ichthyodectid stem-group teleost from the Late Jurassic Solnhofen Limestone of Germany, showing preserved colouration

Ray-finned fish (Actinopterygii) were major components of Jurassic freshwater and marine ecosystems. Archaic "palaeoniscoid" fish, which were common in both marine and freshwater habitats during the preceding Triassic declined during the Jurassic, being largely replaced by more derived actinopterygian lineages.[217] The oldest known Acipenseriformes, the group that contains living sturgeon and paddlefish, are from the Early Jurassic.[218] Amiiform fish (which today only includes the bowfin) first appeared during the Early Jurassic, represented by Caturus from the Pliensbachian of Britain; after their appearance in the western Tethys, they expanded to Africa, North America and Southeast and East Asia by the end of the Jurassic. Pycnodontiformes, which first appeared in the western Tethys during the Late Triassic, expanded to South America and Southeast Asia by the end of the Jurassic, having a high diversity in Europe during the Late Jurassic.[219] During the Jurassic, the Ginglymodi, the only living representatives being gars (Lepisosteidae) were diverse in both freshwater and marine environments. The oldest known representatives of anatomically modern gars appeared during the Upper Jurassic.[220] Stem-group teleosts, which make up over 99% of living Actinopterygii, had first appeared during the Triassic in the western Tethys; they underwent a major diversification beginning in the Late Jurassic, with early representatives of modern teleost clades such as Elopomorpha and Osteoglossoidei appearing during this time.[221][222] The Pachycormiformes, a group of marine stem-teleosts, first appeared in the Early Jurassic and included both tuna-like predatory and filter-feeding forms, the latter included the largest bony fish known to have existed: Leedsichthys, with an estimated maximum length of over 15 metres, known from the late Middle to Late Jurassic.[223]

Chondrichthyes

Fossil of Palaeocarcharias, the oldest known lamniform shark
Fossil of Palaeocarcharias, the oldest known lamniform shark

During the Early Jurassic, the shark-like hybodonts, which represented the dominant group of chondrichthyans during the preceding Triassic, were common in both marine and freshwater settings; however, by the Late Jurassic, hybodonts had become minor components of most marine communities, having been largely replaced by modern neoselachians, but remained common in freshwater and restricted marine environments.[224][225] The Neoselachii, which contains all living sharks and rays, radiated beginning in the Early Jurassic.[226] The oldest known ray (Batoidea) is Antiquaobatis from the Pliensbachian of Germany.[227] Jurassic batoids known from complete remains retain a conservative, guitarfish-like morphology.[228] The oldest known Hexanchiformes and carpet sharks (Orectolobiformes) are from the Early Jurassic (Pliensbachian & Toarcian, respectively) of Europe.[229][230] The oldest known members of the Heterodontiformes, the only living member of which is the bullhead shark (Heterodontus), first appeared in the Early Jurassic, with representatives of the living genus appearing during the Late Jurassic.[231] The oldest known mackerel sharks (Lamniformes) are from the Middle Jurassic, represented by the genus Palaeocarcharias, which has an orectolobiform-like body but shares key similarities in tooth histology with lamniformes, including the absence of orthodentine.[232] The oldest record of angelsharks (Squatiniformes) is Pseudorhina from the Late Jurassic (Oxfordian–Tithonian) of Europe, which already has a bodyform similar to living members of the order.[233] The oldest known remains of Carcharhiniformes, the largest order of living sharks, first appear in the late Middle Jurassic (Bathonian) of the western Tethys (England and Morocco). Known dental and exceptionally preserved body remains of Jurassic Carchariniformes are similar to those of living catsharks.[234] Synechodontiformes, an extinct group of sharks closely related to Neoselachii, were also widespread during the Jurassic.[235] The oldest remains of modern chimaeras are from the Early Jurassic of Europe, with members of the living family Callorhinchidae appearing during the Middle Jurassic. Unlike living chimaeras, these were found in shallow water settings.[236] The closely related Squaloraja and myriacanthoids are also known from the Jurassic of Europe.[237]

Insects and arachnids

Lichnomesopsyche daohugouensis , an extinct mesopsychid scorpionfly from the Late Jurassic of China
Lichnomesopsyche daohugouensis , an extinct mesopsychid scorpionfly from the Late Jurassic of China

There appears to have been no major extinction of insects at the Triassic–Jurassic boundary.[70] Many important insect fossil localities are known from the Jurassic of Eurasia, the most important being the Karabastau Formation of Kazakhstan and the various Yanliao Biota deposits in Inner Mongolia, China, such as the Daohugou Bed, dating to the Callovian–Oxfordian. The diversity of insects stagnated throughout the Early and Middle Jurassic, but during the latter third of the Jurassic origination rates increased substantially while extinction rates remained flat.[238] The increasing diversity of insects in the Middle–Late Jurassic corresponds with a substantial increase in the diversity of insect mouthparts.[239] The Middle to Late Jurassic was a time of major diversification for beetles.[240] Weevils first appear in the fossil record during the Middle to Late Jurassic, but are suspected to have originated during the Late Triassic to Early Jurassic.[241] The oldest known lepidopterans (the group containing butterflies and moths) are known from the Triassic–Jurassic boundary, with wing scales belonging to the suborder Glossata and Micropterigidae-grade moths from the deposits of this age in Germany.[242] Modern representatives of both dragonflies and damselflies also first appeared during the Jurassic.[243] Although modern representatives are not known until the Cenozoic, ectoparasitic insects thought to represent primitive fleas, belonging to the family Pseudopulicidae, are known from the Middle Jurassic of Asia. These insects are substantially different from modern fleas, lacking the specialised morphology of the latter and being larger.[244][245] Parasitoid wasps (Apocrita) first appeared during the Early Jurassic and subsequently became widespread, reshaping terrestrial food webs.[246] The Jurassic saw also saw the first appearances of several other groups of insects, including Phasmatodea (stick insects),[247] Mantophasmatidae,[248] Embioptera (webspinners),[249] and Raphidioptera (snakeflies).[250]

Mongolarachne from the Late Jurassic of China
Mongolarachne from the Late Jurassic of China

Only a handful of records of mites are known from the Jurassic, including Jureremus, an oribatid mite belonging to the family Cymbaeremaeidae known from the Late Jurassic of Britain and Russia,[251] and a member of the still living orbatid genus Hydrozetes from the Early Jurassic of Sweden.[252] Spiders diversified through the Jurassic.[253] The Early Jurassic Seppo koponeni may represent a stem group to Palpimanoidea.[254] Eoplectreurys from the Middle Jurassic of China is considered a stem lineage of Synspermiata. The oldest member of the family Archaeidae, Patarchaea, is known from the Middle Jurassic of China.[253] Mongolarachne from the Middle Jurassic of China is among the largest known fossil spiders, with legs over 5 centimetres long.[255] The only scorpion known from the Jurassic is Liassoscorpionides from the Early Jurassic of Germany, of uncertain placement.[256] Eupnoi harvestmen (Opiliones) are known from the Middle Jurassic of China, including members of the family Sclerosomatidae.[257][258]

Marine invertebrates

End-Triassic extinction

During the end-Triassic extinction, 46%–72% of all marine genera became extinct. The effects of the end Triassic extinction were greatest at tropical latitudes and were more severe in Panthalassa than the Tethys or Boreal oceans. Tropical reef ecosystems collapsed during the event, and would not fully recover until much later in the Jurassic. Sessile filter feeders and photosymbiotic organisms were among most severely affected.[259]

Marine ecosystems

Having declined at the Triassic–Jurassic boundary, reefs substantially expanded during the Late Jurassic, including both sponge reefs and scleractinian coral reefs. Late Jurassic reefs were similar in form to modern reefs but had more microbial carbonates and hypercalcified sponges, and had weak biogenic binding. Reefs sharply declined at the close of the Jurassic,[260] which caused an associated drop in diversity in decapod crustaceans.[261] The earliest planktonic foraminifera, which constitute the suborder Globigerinina, are known from the late Early Jurassic (mid-Toarcian) of the western Tethys, expanding across the whole Tethys by the Middle Jurassic and becoming globally distributed in tropical latitudes by the Late Jurassic.[262] Coccolithophores and dinoflagellates, which had first appeared during the Triassic, radiated during the Early to Middle Jurassic, becoming prominent members of the phytoplankton.[263] Microconchid tube worms, the last remaining order of Tentaculita, a group of animals of uncertain affinities that were convergent on Spirorbis tube worms, were rare after the Triassic and had become reduced to the single genus Punctaconchus, which became extinct in the late Bathonian.[264] The oldest known diatom is from Late Jurassic–aged amber from Thailand, assigned to the living genus Hemiaulus.[265]

Echinoderms

Crinoids diversified throughout the Jurassic, reaching their peak Mesozoic diversity during the Late Jurassic, primarily due to the radiation of sessile forms belonging to the orders Cyrtocrinida and Millericrinida.[266] Echinoids (sea urchins) underwent substantial diversification beginning in the Early Jurassic, primarily driven by the radiation of irregular (asymmetrical) forms, which were adapting to deposit feeding. Rates of diversification sharply dropped during the Late Jurassic.[267]

Crustaceans

Eryon, a polychelidan decapod crustacean from the Late Jurassic of Germany.
Eryon, a polychelidan decapod crustacean from the Late Jurassic of Germany.

The Jurassic was a significant time for the evolution of decapods.[261] The first true crabs (Brachyura) are known from the Early Jurassic, with the earliest being Eocarcinus praecursor from the early Pliensbachian of England, which lacked the crab-like morphology (carcinisation) of modern crabs,[268] and Eoprosopon klugi from the late Pliensbachian of Germany, which may belong to the living family Homolodromiidae.[269] Most Jurassic crabs are known only from carapace pieces, which makes it difficult to determine their relationships.[270] While rare in the Early and Middle Jurassic, crabs became abundant during the Late Jurassic as they expanded from their ancestral silty sea floor habitat into hard substrate habitats like reefs, with crevices in reefs providing refuge from predators.[270][261] Hermit crabs also first appeared during the Jurassic, with the earliest known being Schobertella hoelderi from the late Hettangian of Germany.[271] Early hermit crabs are associated with ammonite shells rather than those of gastropods.[272] Glypheids, which today are only known from two species, reached their peak diversity during the Jurassic, with around 150 species out of a total fossil record of 250 known from the period.[273] Jurassic barnacles were of low diversity compared to present,[274] but several important evolutionary innovations are known, including the first appearances of calcite shelled forms and species with an epiplanktonic mode of life.[275]

Brachiopods

Brachiopod diversity declined during the Triassic–Jurassic extinction. Spire-bearing brachiopods (Spiriferinida and Athyridida) did not recover their biodiversity, becoming extinct in the TOAE.[276] Rhynchonellida and Terebratulida also declined during the Triassic–Jurassic extinction but rebounded during the Early Jurassic; neither clade underwent much morphological variation.[277] Brachiopods substantially declined in the Late Jurassic; the causes are poorly understood. Proposed reasons include increased predation, competition with bivalves, enhanced bioturbation or increased grazing pressure.[278]

Bryozoans

Like the preceding Triassic, bryozoan diversity was relatively low compared to the Paleozoic. The vast majority of Jurassic bryozoans are members of Cyclostomatida, which experienced a radiation during the Middle Jurassic, with all Jurassic representatives belonging to the suborders Tubuliporina and Cerioporina. Cheilostomata, the dominant group of modern bryozoans, first appeared during the Late Jurassic.[279]

Molluscs

Bivalves

The end-Triassic extinction had a severe impact on bivalve diversity, though it had little impact on bivalve ecological diversity. The extinction was selective, having less of an impact on deep burrowers, but there is no evidence of a differential impact between surface-living (epifaunal) and burrowing (infaunal) bivalves.[280] Bivalve family level diversity after the Early Jurassic was static, though genus diversity experienced a gradual increase throughout the period.[281] Rudists, the dominant reef-building organisms of the Cretaceous, first appeared in the Late Jurassic (mid-Oxfordian) in the northern margin of the western Tethys, expanding to the eastern Tethys by the end of the Jurassic.[282]

Cephalopods
Fossil specimen of Proteroctopus from the Middle Jurassic of France, formerly thought to be worlds oldest known octopus
Fossil specimen of Proteroctopus from the Middle Jurassic of France, formerly thought to be worlds oldest known octopus

Ammonites were devastated by the end-Triassic extinction, with only a handful of genera belonging to the family Psiloceratidae of the suborder Phylloceratina surviving and becoming ancestral to all later Jurassic and Cretaceous ammonites. Ammonites explosively diversified during the Early Jurassic, with the orders Psiloceratina, Ammonitina, Lytoceratina, Haploceratina, Perisphinctina and Ancyloceratina all appearing during the Jurassic. Ammonite faunas during the Jurassic were regional, being divided into around 20 distinguishable provinces and subprovinces in two realms, the northern high latitude Pan-Boreal realm, consisting of the Arctic, northern Panthalassa and northern Atlantic regions, and the equatorial–southern Pan-Tethyan realm, which included the Tethys and most of Panthalassa.[283]

The oldest definitive records of the squid-like belemnites are from the earliest Jurassic (Hettangian–Sinemurian) of Europe and Japan; they expanded worldwide during the Jurassic.[284] Belemnites were shallow-water dwellers, inhabiting the upper 200 metres of the water column on the continental shelves and in the littoral zone. They were key components of Jurassic ecosystems, both as predators and prey, as evidenced by the abundance of belemnite guards in Jurassic rocks.[285]

The earliest vampyromorphs, of which the only living member is the vampire squid, first appeared during the Early Jurassic.[286] The earliest octopuses appeared during the Middle Jurassic, having split from their closest living relatives, the vampyromorphs, during the Triassic to Early Jurassic.[287] All Jurassic octopuses are solely known from the hard gladius.[287][288] Octopuses likely originated from bottom-dwelling (benthic) ancestors which lived in shallow environments.[287] Proteroctopus from the late Middle Jurassic La Voulte-sur-Rhône lagerstätte, previously interpreted as an early octopus, is now thought to be a basal taxon outside the clade containing vampyromorphs and octopuses.[289]

Discover more about Fauna related topics

Holotype

Holotype

A holotype is a single physical example of an organism, known to have been used when the species was formally described. It is either the single such physical example or one of several examples, but explicitly designated as the holotype. Under the International Code of Zoological Nomenclature (ICZN), a holotype is one of several kinds of name-bearing types. In the International Code of Nomenclature for algae, fungi, and plants (ICN) and ICZN, the definitions of types are similar in intent but not identical in terminology or underlying concept.

Platysuchus

Platysuchus

Platysuchus is an extinct genus of teleosaurid crocodyliform from the Early Jurassic (Toarcian) of southern Germany and Luxembourg.

Crocodylomorpha

Crocodylomorpha

Crocodylomorpha is a group of pseudosuchian archosaurs that includes the crocodilians and their extinct relatives. They were the only members of Pseudosuchia to survive the end-Triassic extinction.

Aetosaur

Aetosaur

Aetosaurs are heavily armored reptiles belonging to the extinct order Aetosauria. They were medium- to large-sized omnivorous or herbivorous pseudosuchians, part of the branch of archosaurs more closely related to crocodilians than to birds and other dinosaurs. All known aetosaurs are restricted to the Late Triassic, and in some strata from this time they are among the most abundant fossil vertebrates. They have small heads, upturned snouts, erect limbs, and a body ornamented with four rows of plate-like osteoderms. Aetosaur fossil remains are known from Europe, North and South America, parts of Africa, and India. Since their armoured plates are often preserved and are abundant in certain localities, aetosaurs serve as important Late Triassic tetrapod index fossils. Many aetosaurs had wide geographic ranges, but their stratigraphic ranges were relatively short. Therefore, the presence of particular aetosaurs can accurately date a site in which they are found.

Ecological niche

Ecological niche

In ecology, a niche is the match of a species to a specific environmental condition. It describes how an organism or population responds to the distribution of resources and competitors and how it in turn alters those same factors. "The type and number of variables comprising the dimensions of an environmental niche vary from one species to another [and] the relative importance of particular environmental variables for a species may vary according to the geographic and biotic contexts".

Crocodilia

Crocodilia

Crocodilia is an order of mostly large, predatory, semiaquatic reptiles, known as crocodilians. They first appeared 95 million years ago in the Late Cretaceous period and are the closest living relatives of birds, as the two groups are the only known survivors of the Archosauria. Members of the order's total group, the clade Pseudosuchia, appeared about 250 million years ago in the Early Triassic period, and diversified during the Mesozoic era. The order Crocodilia includes the true crocodiles, the alligators and caimans, and the gharial and false gharial. Although the term 'crocodiles' is sometimes used to refer to all of these, crocodilians is a less ambiguous vernacular term for members of this group.

Metriorhynchidae

Metriorhynchidae

Metriorhynchidae is an extinct family of specialized, aquatic metriorhynchoid crocodyliforms from the Middle Jurassic to the Early Cretaceous period of Europe, North America and South America. The name Metriorhynchidae was coined by the Austrian zoologist Leopold Fitzinger in 1843. The group contains two subfamilies, the Metriorhynchinae and the Geosaurinae. They represent the most marine adapted of all archosaurs.

Crocodyliformes

Crocodyliformes

Crocodyliformes is a clade of crurotarsan archosaurs, the group often traditionally referred to as "crocodilians". They are the first members of Crocodylomorpha to possess many of the features that define later relatives. They are the only pseudosuchians to survive the K-Pg extinction event.

Neosuchia

Neosuchia

Neosuchia is a clade within Mesoeucrocodylia that includes all modern extant crocodilians and their closest fossil relatives. It is defined as the most inclusive clade containing all crocodylomorphs more closely related to Crocodylus niloticus than to Notosuchus terrestris. Members of Neosuchia generally share a crocodilian-like bodyform adapted to freshwater aquatic life, as opposed to the terrestrial habits of more basal crocodylomorph groups. The earliest neosuchian is suggested to be the Early Jurassic Calsoyasuchus, which lived during the Sinemurian and Pliensbachian stages in North America. It is often identified as a member of Goniopholididae, though this is disputed, and the taxon may lie outside Neosuchia, which places the earliest records of the group in the Middle Jurassic.

Calsoyasuchus

Calsoyasuchus

Calsoyasuchus is a genus of crocodylomorph that lived in the Early Jurassic. Its fossilized remains were found in the Sinemurian-Pliensbachian-age Kayenta Formation on Navajo Nation land in Coconino County, Arizona, United States. Formally described as C. valliceps, it is known from a single incomplete skull which is unusually derived for such an early crocodile relative. This genus was described in 2002 by Ronald Tykoski and colleagues; the specific name means "valley head" and refers to a deep groove along the midline of the nasal bones and frontal bones. It has often been interpreted as the earliest diverging member of Goniopholididae, but other studies have recovered it in various other positions.

Goniopholididae

Goniopholididae

Goniopholididae is an extinct family of moderate-sized semi-aquatic neosuchian crocodyliformes. Their bodyplan and morphology are convergent on living crocodilians. They lived across Laurasia between the Middle Jurassic and the Late Cretaceous.

Notosuchia

Notosuchia

Notosuchia is a suborder of primarily Gondwanan mesoeucrocodylian crocodylomorphs that lived during the Jurassic and Cretaceous. Some phylogenies recover Sebecosuchia as a clade within Notosuchia, others as a sister group ; if Sebecosuchia is included within Notosuchia its existence is pushed into the Middle Miocene, about 11 million years ago. Fossils have been found from South America, Africa, Asia, and Europe. Notosuchia was a clade of terrestrial crocodilians that evolved a range of feeding behaviours, including herbivory (Chimaerasuchus), omnivory (Simosuchus), and terrestrial hypercarnivory (Baurusuchus). It included many members with highly derived traits unusual for crocodylomorphs, including mammal-like teeth, flexible bands of shield-like body armor similar to those of armadillos (Armadillosuchus), and possibly fleshy cheeks and pig-like snouts (Notosuchus). The suborder was first named in 1971 by Zulma Gasparini and has since undergone many phylogenetic revisions.

Source: "Jurassic", Wikipedia, Wikimedia Foundation, (2023, March 17th), https://en.wikipedia.org/wiki/Jurassic.

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