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Helix (gastropod)

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Helix
Temporal range: Oligocene–recent
Helix pomatia 89a.jpg
Helix pomatia
Scientific classification e
Kingdom: Animalia
Phylum: Mollusca
Class: Gastropoda
Subclass: Heterobranchia
Order: Stylommatophora
Family: Helicidae
Subfamily: Helicinae
Tribe: Helicinini
Genus: Helix
Linnaeus, 1758
Type species
Helix pomatia

Helix is a genus of large, air-breathing land snails native to Europe and the Mediterranean region. They are the type genus of the family Helicidae, and are among the first animal genera described by Carl Linnaeus.[1] Members of the genus first appeared during the Oligocene.[2] Like most land snails, Helix species are hermaphroditic, and like other helicids, a courting pair will impale each other with hormone-rich love darts prior to exchanging sperm. Well-known species include Helix pomatia (Roman snail, Burgundy snail, or edible snail) and Helix lucorum (Turkish snail). Cornu aspersum (garden snail) – another well-known helicid – was originally described as a member of Helix (as "Helix aspersa"), but the prevailing classification places it as the sole member of the sister genus Cornu.[1]

Helix snails have been harvested for human consumption since prehistoric times.[3] In the common era, members of the genus (particularly H. pomatia) are farmed throughout the world for the dish escargot, an hors d'oeuvre. The establishment of snail farms outside of Europe has introduced several species to North America, South America, and Africa, where some escapees have established themselves as invasive species.[4][5][6][7]

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Genus

Genus

Genus is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nomenclature, the genus name forms the first part of the binomial species name for each species within the genus.E.g. Panthera leo (lion) and Panthera onca (jaguar) are two species within the genus Panthera. Panthera is a genus within the family Felidae.

Europe

Europe

Europe is a continent comprising the westernmost peninsulas of Eurasia, located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It shares the continental landmass of Afro-Eurasia with both Africa and Asia. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east. Europe is commonly considered to be separated from Asia by the watershed of the Ural Mountains, the Ural River, the Caspian Sea, the Greater Caucasus, the Black Sea and the waterways of the Turkish Straits.

Helicidae

Helicidae

Helicidae is a large, diverse family of small to large, air-breathing land snails, sometimes called the "typical snails."

Carl Linnaeus

Carl Linnaeus

Carl Linnaeus, also known after his ennoblement in 1761 as Carl von Linné, was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the modern system of naming organisms. He is known as the "father of modern taxonomy". Many of his writings were in Latin; his name is rendered in Latin as Carolus Linnæus and, after his 1761 ennoblement, as Carolus a Linné.

Courtship

Courtship

Courtship is the period wherein some couples get to know each other prior to a possible marriage or committed romantic, de facto relationship. Courtship traditionally may begin after a betrothal and may conclude with the celebration of marriage. A courtship may be an informal and private matter between two people or may be a public affair, or a formal arrangement with family approval. Traditionally, in the case of a formal engagement, it is the role of a male to actively "court" or "woo" a female, thus encouraging her to understand him and her receptiveness to a marriage proposal.

Hormone

Hormone

A hormone is a class of signaling molecules in multicellular organisms that are sent to distant organs by complex biological processes to regulate physiology and behavior. Hormones are required for the correct development of animals, plants and fungi. Due to the broad definition of a hormone, numerous kinds of molecules can be classified as hormones. Among the substances that can be considered hormones, are eicosanoids, steroids, amino acid derivatives, protein or peptides, and gases.

Helix pomatia

Helix pomatia

Helix pomatia, common names the Roman snail, Burgundy snail, or escargot, is a species of large, edible, air-breathing land snail, a pulmonate gastropod terrestrial mollusc in the family Helicidae. It is one of Europe's biggest species of land snail.

Helix lucorum

Helix lucorum

Helix lucorum is a species of large, edible, air-breathing land snail, a terrestrial pulmonate gastropod mollusk in the family Helicidae, the typical snails.

Cornu aspersum

Cornu aspersum

Cornu aspersum, known by the common name garden snail, is a species of land snail in the family Helicidae, which includes some of the most familiar land snails. Of all terrestrial molluscs, this species may well be the most widely known. It was classified under the name Helix aspersa for over two centuries, but the prevailing classification now places it in the genus Cornu.

Heliciculture

Heliciculture

Heliciculture, commonly known as snail farming, is the process of raising edible land snails, primarily for human consumption or cosmetic use. The meat and snail eggs a.k.a. white caviar can be consumed as escargot and as a type of caviar, respectively.

Escargot

Escargot

Snails are considered edible in certain areas such as the Mediterranean region, Africa, or Southeast Asia, while in other cultures it is considered as taboo food. In American English, edible land snails are also called escargot, taken from the French word for 'snail', and the production of snails for consumption is called snail farming or heliciculture. Snails as food date back to ancient times, and it was a dish that was already served on the tables of Ancient Greece and Rome.

Hors d'oeuvre

Hors d'oeuvre

An hors d'oeuvre, appetiser or starter is a small dish served before a meal in European cuisine. Some hors d'oeuvres are served cold, others hot. Hors d'oeuvres may be served at the dinner table as a part of the meal, or they may be served before seating, such as at a reception or cocktail party. Formerly, hors d'oeuvres were also served between courses. There are two types of hors d'oeuvre from service point of view:General hors d'oeuvre Classical hors d'oeuvre

Taxonomy

Some taxonomists remove the species "Helix aperta", "Helix aspersa", and "Helix mazzullii" from the genus Helix and place them in their own monotypic genera as Cantareus apertus, Cornu aspersum,[8] and Cantareus mazzullii.

At the beginning in the mid-1700s, the generic name Helix had been used for almost all terrestrial gastropods; later this was restricted to species with helicoid habitus, including zonitids and other groups. In the course of the 1800s, more groups were removed, but prior to 1900, several thousand helicid and hygromiid species of Europe and abroad had still been classified in the genus Helix.[9][10] By the early 1900s, the genus was split into many separate genera, leaving only some 30 species closely related to its type species Helix pomatia in the genus.

The genus Helix has many synonyms:

  • Callunea Scudder, 1882
  • Cochlea Da Costa, 1778
  • Coenatoria Held, 1838
  • Cunula Pallary, 1936
  • Glischrus S. Studer, 1820
  • Glischrus (Helix) Linnaeus, 1758
  • Helicites W. Martin, 1809 (Established for fossils of Helix to distinguish them from extant members of that taxon. Invalid: available only for the purposes of the Principle of Homonymy (Art. 20))
  • Helicogena A. Férussac, 1821
  • Helix (Helix) Linnaeus, 1758· accepted, alternate representation
  • Helix (Pelasga) P. Hesse, 1908· accepted, alternate representation
  • Megastoma Scudder, 1882
  • Naegelea P. Hesse, 1918
  • Pachyphallus P. Hesse, 1918
  • Pentataenia A. Schmidt, 1855 (junior objective synonym)
  • Physospira C.R. Boettger, 1914
  • Pomatia Beck, 1837
  • Pomatiana Fagot, 1903
  • Pomatiella Pallary, 1909
  • Pseudofigulina P. Hesse, 1917
  • Rhododerma P. Hesse, 1918
  • Tacheopsis Boettger, 1909
  • Tammouzia Pallary, 1939

Many species of Helix have been brought into synonymy:

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Cantareus apertus

Cantareus apertus

Cantareus apertus, commonly known as the green garden snail, is a species of air-breathing land snail, a terrestrial pulmonate gastropod mollusc in the family Helicidae, the typical snails.

Carl Linnaeus

Carl Linnaeus

Carl Linnaeus, also known after his ennoblement in 1761 as Carl von Linné, was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the modern system of naming organisms. He is known as the "father of modern taxonomy". Many of his writings were in Latin; his name is rendered in Latin as Carolus Linnæus and, after his 1761 ennoblement, as Carolus a Linné.

10th edition of Systema Naturae

10th edition of Systema Naturae

The 10th edition of Systema Naturae is a book written by Swedish naturalist Carl Linnaeus and published in two volumes in 1758 and 1759, which marks the starting point of zoological nomenclature. In it, Linnaeus introduced binomial nomenclature for animals, something he had already done for plants in his 1753 publication of Species Plantarum.

Chilostoma achates

Chilostoma achates

Chilostoma achates is a species of medium-sized, air-breathing, land snail, a terrestrial pulmonate gastropod mollusk in the family Helicidae, the true snails. IUCN has defined the snail as Near threatened, due to the development of tourism.

Fossarus ambiguus

Fossarus ambiguus

Fossarus ambiguus is a species of sea snail, a marine gastropod mollusk in the family Planaxidae.

Pila ampullacea

Pila ampullacea

Pila ampullacea, is a species of freshwater snail with an operculum, an aquatic gastropod mollusk in the family Ampullariidae, the apple snails.

Chilostoma cingulatum

Chilostoma cingulatum

Chilostoma cingulatum is a species of medium-sized, air-breathing land snail, a terrestrial pulmonate gastropod mollusk in the family Helicidae, the true snails.SubspeciesChilostoma cingulatum adamii (Pini, 1876) Chilostoma cingulatum alzonai K. L. Pfeiffer, 1951 Chilostoma cingulatum anauniense (De Betta, 1852) Chilostoma cingulatum anconae (Gentiluomo, 1868) Chilostoma cingulatum appelii (Kobelt, 1876) Chilostoma cingulatum asperulum (Ehrmann, 1910) Chilostoma cingulatum baldense (Rossmässler, 1839) Chilostoma cingulatum bizona (Rossmässler, 1842) Chilostoma cingulatum boccavallense K. L. Pfeiffer, 1951 Chilostoma cingulatum carrarense (Strobel, 1852) Chilostoma cingulatum cingulatum (S. Studer, 1820) Chilostoma cingulatum colubrinum (De Cristofori & Jan, 1832) Chilostoma cingulatum frigidescens (Del Prete, 1879) Chilostoma cingulatum frigidissimum (Paulucci, 1881) Chilostoma cingulatum frigidosum (Pollonera, 1890) Chilostoma cingulatum gobanzi (Frauenfeld, 1867) Chilostoma cingulatum hermesianum (Pini, 1874) Chilostoma cingulatum infernale (P. Hesse, 1931) Chilostoma cingulatum insubricum (De Cristofori & Jan, 1832) Chilostoma cingulatum medoacense (Adami, 1886) Chilostoma cingulatum montanum (Paulucci, 1881) Chilostoma cingulatum nicatis (Costa, 1836) Chilostoma cingulatum nicolisianum (Adami, 1886) Chilostoma cingulatum peregrini Falkner, 1998 Chilostoma cingulatum philippii (Kobelt, 1905) Chilostoma cingulatum preslii (Rossmässler, 1836) Chilostoma cingulatum sentinense (Piersanti, 1833) Chilostoma cingulatum transiens (Adami, 1886)

Hippeutis complanatus

Hippeutis complanatus

Hippeutis complanatus, or the flat ram's-horn snail, is a species of minute air-breathing freshwater snail, an aquatic pulmonate gastropod mollusk or micromollusk in the family Planorbidae, the ram's horn snails.

Bathyomphalus contortus

Bathyomphalus contortus

Bathyomphalus contortus is a species of small air-breathing freshwater snail, an aquatic pulmonate gastropod mollusk in the family Planorbidae, the ram's horn snails and their allies.

Marisa cornuarietis

Marisa cornuarietis

Marisa cornuarietis, common name the Colombian ramshorn apple snail, is a species of large freshwater snail with an operculum, an aquatic gastropod mollusc in the family Ampullariidae, the apple snail family.

Amphibola crenata

Amphibola crenata

Amphibola crenata is a species of air-breathing snail with an operculum, a pulmonate gastropod mollusc which lives in a habitat that is intermediate between the land and the sea, not entirely terrestrial and not entirely marine. This is not a true land snail, but it is also not a true sea snail. Unlike almost all other snails that have opercula, this species breathes air. It is common in New Zealand.

George Montagu (naturalist)

George Montagu (naturalist)

George Montagu was an English army officer and naturalist. He was known for his pioneering Ornithological Dictionary of 1802, which for the first time accurately defined the status of Britain's birds. He is remembered today for species such as the Montagu's harrier, named for him.

Description

An unidentified Helix species from Southern Europe
An unidentified Helix species from Southern Europe

In addition to the hard, calcareous shell that covers and protects the internal organs, the head and foot regions can be observed when the snails are fully extended. When they are active, the organs such as the lung, heart, kidney, and intestines remain inside the shell; only the head and foot emerge.

The head of the snail has two pairs of tentacles; the upper and larger pair contains the eyes, and the lower pair is used to feel the ground in front. The mouth is located just underneath the head. The tentacles can be withdrawn or extended depending on the situation. The mouth has a tongue called a "radula" that is composed of many fine, chitinous teeth. This serves for rasping and cutting food.

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Lung

Lung

The lungs are the primary organs of the respiratory system in humans and most other animals, including some snails and a small number of fish. In mammals and most other vertebrates, two lungs are located near the backbone on either side of the heart. Their function in the respiratory system is to extract oxygen from the air and transfer it into the bloodstream, and to release carbon dioxide from the bloodstream into the atmosphere, in a process of gas exchange. The pleurae, which are thin, smooth, and moist, serve to reduce friction between the lungs and chest wall during breathing, allowing for easy and effortless movements of the lungs.

Heart

Heart

The heart is a muscular organ in most animals. This organ pumps blood through the blood vessels of the circulatory system. The pumped blood carries oxygen and nutrients to the body, while carrying metabolic waste such as carbon dioxide to the lungs. In humans, the heart is approximately the size of a closed fist and is located between the lungs, in the middle compartment of the chest.

Kidney

Kidney

The kidneys are two reddish-brown bean-shaped organs found in vertebrates. They are located on the left and right in the retroperitoneal space, and in adult humans are about 12 centimetres in length. They receive blood from the paired renal arteries; blood exits into the paired renal veins. Each kidney is attached to a ureter, a tube that carries excreted urine to the bladder.

Tongue

Tongue

The tongue is a muscular organ in the mouth of a typical tetrapod. It manipulates food for chewing and swallowing as part of the digestive process, and is the primary organ of taste. The tongue's upper surface (dorsum) is covered by taste buds housed in numerous lingual papillae. It is sensitive and kept moist by saliva and is richly supplied with nerves and blood vessels. The tongue also serves as a natural means of cleaning the teeth. A major function of the tongue is the enabling of speech in humans and vocalization in other animals.

Radula

Radula

The radula is an anatomical structure used by mollusks for feeding, sometimes compared to a tongue. It is a minutely toothed, chitinous ribbon, which is typically used for scraping or cutting food before the food enters the esophagus. The radula is unique to the mollusks, and is found in every class of mollusk except the bivalves, which instead use cilia, waving filaments that bring minute organisms to the mouth.

Behaviour

From April through the northern summer, the number of snails copulating increases due to the higher temperature and humidity, which enhance the possibility of oviposition. The pulmonate snails are hermaphroditic, meaning that both female and male sexual organs are present in the same individual. The snails produce both eggs and sperm in the ovotestis (also called the hermaphrodite gland), but it is later separated into two divisions, a sperm duct and oviduct, respectively.

Mating takes several hours, sometimes a day. H. aspersa snails stab a calcite spine, known as a "love dart", at their partner. The love dart is coated with a mucus that contains a chemical that enables more than twice as many sperm to survive inside the recipient. A few days after mating, the eggs are laid in the soil. The eggs are usually 4–6 mm in diameter.

After snails hatch from the egg, they mature in one or more years, depending on where the organism lives. Maturity takes two years in Southern California, while it takes only 10 months in South Africa.

The size of the adult snails slightly varies with species. H. aspersa grows up to 35 mm in height and width, whereas H. pomatia grows up to 45 mm. The lifespan of snails in the wild is typically 2-3 years.

Some snails may live longer, perhaps even 30 years or older in the case of the Roman snail[11] but most live less than 8 years. Many deaths are due to predators and parasites.

Since its publication in March 1974 issue of Natural History[12] an estimate for the highest speed of garden snail equal to 0.03 mph (1.3 cm/s) became popular.[13] However, the accuracy of this estimate has been questioned by Robert Cameron, who pointed out that in competitions between snails, only 2.4 mm/s speed had been achieved.[14] There is a need for clarification and behavioural observation here though: competitions are usually held on the flat, whereas a snail's natural preference is vertical ascent, possibly an instinct derived from hatching in an earth "pod" and having to climb upwards to the surface.

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Hermaphrodite

Hermaphrodite

In reproductive biology, a hermaphrodite is an organism that has both kinds of reproductive organs and can produce both gametes associated with male and female sexes.

Sperm

Sperm

Sperm is the male reproductive cell, or gamete, in anisogamous forms of sexual reproduction. Animals produce motile sperm with a tail known as a flagellum, which are known as spermatozoa, while some red algae and fungi produce non-motile sperm cells, known as spermatia. Flowering plants contain non-motile sperm inside pollen, while some more basal plants like ferns and some gymnosperms have motile sperm.

Oviduct

Oviduct

The oviduct in mammals, is the passageway from an ovary. In human females this is more usually known as the Fallopian tube or uterine tube. The eggs travel along the oviduct. These eggs will either be fertilized by spermatozoa to become a zygote, or will degenerate in the body. Normally, these are paired structures, but in birds and some cartilaginous fishes, one or the other side fails to develop, and only one functional oviduct can be found.

Mating

Mating

In biology, mating is the pairing of either opposite-sex or hermaphroditic organisms for the purposes of sexual reproduction. Fertilization is the fusion of two gametes. Copulation is the union of the sex organs of two sexually reproducing animals for insemination and subsequent internal fertilization. Mating may also lead to external fertilization, as seen in amphibians, fishes and plants. For most species, mating is between two individuals of opposite sexes. However, for some hermaphroditic species, copulation is not required because the parent organism is capable of self-fertilization (autogamy); for example, banana slugs.

Love dart

Love dart

A love dart is a sharp, calcareous or chitinous dart which some hermaphroditic land snails and slugs create. Love darts are both formed and stored internally in a dart sac. These darts are made in sexually mature animals only, and are used as part of the sequence of events during courtship, before actual mating takes place. Darts are quite large compared to the size of the animal: in the case of the semi-slug genus Parmarion, the length of a dart can be up to one fifth that of the semi-slug's foot.

California

California

California is a state in the Western United States, located along the Pacific Coast. With nearly 39.2 million residents across a total area of approximately 163,696 square miles (423,970 km2), it is the most populous U.S. state and the third-largest by area. It is also the most populated subnational entity in North America and the 34th most populous in the world. The Greater Los Angeles and San Francisco Bay areas are the nation's second and fifth most populous urban regions respectively, with the former having more than 18.7 million residents and the latter having over 9.6 million. Sacramento is the state's capital, while Los Angeles is the most populous city in the state and the second most populous city in the country. San Francisco is the second most densely populated major city in the country. Los Angeles County is the country's most populous, while San Bernardino County is the largest county by area in the country. California borders Oregon to the north, Nevada and Arizona to the east, the Mexican state of Baja California to the south; and it has a coastline along the Pacific Ocean to the west.

South Africa

South Africa

South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. It is bounded to the south by 2,798 kilometres (1,739 mi) of coastline that stretches along the South Atlantic and Indian Oceans; to the north by the neighbouring countries of Namibia, Botswana, and Zimbabwe; and to the east and northeast by Mozambique and Eswatini. It also completely enclaves the country Lesotho. It is the southernmost country on the mainland of the Old World, and the second-most populous country located entirely south of the equator, after Tanzania. South Africa is a biodiversity hotspot, with unique biomes, plant and animal life. With over 60 million people, the country is the world's 24th-most populous nation and covers an area of 1,221,037 square kilometres. Pretoria is the administrative capital, while Cape Town, as the seat of Parliament, is the legislative capital. Bloemfontein has traditionally been regarded as the judicial capital. The largest city, and site of highest court is Johannesburg.

Natural History (magazine)

Natural History (magazine)

Natural History is a natural history magazine published in the United States. The stated mission of the magazine is to promote public understanding and appreciation of nature and science.

Miles per hour

Miles per hour

Miles per hour is a British imperial and United States customary unit of speed expressing the number of miles travelled in one hour. It is used in the United Kingdom, the United States, and a number of smaller countries, most of which are UK or US territories, or have close historical ties with the UK or US.

Respiration

Burgundy snail (H. pomatia)
Burgundy snail (H. pomatia)

Since snails in the genus Helix are terrestrial rather than freshwater or marine, they have developed a simple lung for respiration. (Most other snails and gastropods have gills, instead.)

Oxygen is carried by the blood pigment hemocyanin. Both oxygen and carbon dioxide diffuse in and out of blood through the capillaries. A muscular valve regulates the process of opening and closing the entrance of the lung. When the valve opens, the air can either enter or leave the lung. The valve plays an important role in reducing water loss and preventing drowning.

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Gill

Gill

A gill is a respiratory organ that many aquatic organisms use to extract dissolved oxygen from water and to excrete carbon dioxide. The gills of some species, such as hermit crabs, have adapted to allow respiration on land provided they are kept moist. The microscopic structure of a gill presents a large surface area to the external environment. Branchia is the zoologists' name for gills.

Oxygen

Oxygen

Oxygen is the chemical element with the symbol O and atomic number 8. It is a member of the chalcogen group in the periodic table, a highly reactive nonmetal, and an oxidizing agent that readily forms oxides with most elements as well as with other compounds. Oxygen is Earth's most abundant element, and after hydrogen and helium, it is the third-most abundant element in the universe. At standard temperature and pressure, two atoms of the element bind to form dioxygen, a colorless and odorless diatomic gas with the formula O2. Diatomic oxygen gas currently constitutes 20.95% of the Earth's atmosphere, though this has changed considerably over long periods of time. Oxygen makes up almost half of the Earth's crust in the form of oxides.

Hemocyanin

Hemocyanin

Hemocyanins (also spelled haemocyanins and abbreviated Hc) are proteins that transport oxygen throughout the bodies of some invertebrate animals. These metalloproteins contain two copper atoms that reversibly bind a single oxygen molecule (O2). They are second only to hemoglobin in frequency of use as an oxygen transport molecule. Unlike the hemoglobin in red blood cells found in vertebrates, hemocyanins are not confined in blood cells but are instead suspended directly in the hemolymph. Oxygenation causes a color change between the colorless Cu(I) deoxygenated form and the blue Cu(II) oxygenated form.

Carbon dioxide

Carbon dioxide

Carbon dioxide is a chemical compound made up of molecules that each have one carbon atom covalently double bonded to two oxygen atoms. It is found in the gas state at room temperature, and as the source of available carbon in the carbon cycle, atmospheric CO2 is the primary carbon source for life on Earth. In the air, carbon dioxide is transparent to visible light but absorbs infrared radiation, acting as a greenhouse gas. Carbon dioxide is soluble in water and is found in groundwater, lakes, ice caps, and seawater. When carbon dioxide dissolves in water, it forms carbonate and mainly bicarbonate, which causes ocean acidification as atmospheric CO2 levels increase.

Ecology

Helix snails prefer cool, damp environments, as they easily suffer moisture loss. Snails are most active at night and after rainfall. During unfavourable conditions, a snail remains inside its shell, usually under rocks or other hiding places, to avoid being discovered by predators. In dry climates, snails naturally congregate near water sources, including artificial sources such as wastewater outlets of air conditioners.

The common garden snail (H.aspersa) is herbivorous. These snails are able to digest most vegetation, including carrots and lettuce. They also have a specialized crop of symbiotic bacteria that aid in their digestion, especially with the breakdown of the polysaccharide cellulose into simple sugars.

Many predators, both specialist and generalist, feed on snails. Some animals, such as the song thrush, break the shell of the snail by hammering it against a hard object, such as stone, to expose its edible insides. Other predators, such as some species of frogs, circumvent the need to break snail shells by simply swallowing the snail whole, shell and all.

Some carnivorous species of snails, such as the decollate snail and the rosy wolf snail, also prey on Helix snails. Such carnivorous snails are commercially grown and sold to combat pest snail species. Many of these also escape into the wild, where they prey on indigenous snails, such as the Cuban land snails of the genus Polymita, and the indigenous snails of Hawaii.

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Herbivore

Herbivore

A herbivore is an animal anatomically and physiologically adapted to eating plant material, for example foliage or marine algae, for the main component of its diet. As a result of their plant diet, herbivorous animals typically have mouthparts adapted to rasping or grinding. Horses and other herbivores have wide flat teeth that are adapted to grinding grass, tree bark, and other tough plant material.

Carrot

Carrot

The carrot is a root vegetable, typically orange in color, though purple, black, red, white, and yellow cultivars exist, all of which are domesticated forms of the wild carrot, Daucus carota, native to Europe and Southwestern Asia. The plant probably originated in Persia and was originally cultivated for its leaves and seeds. The most commonly eaten part of the plant is the taproot, although the stems and leaves are also eaten. The domestic carrot has been selectively bred for its enlarged, more palatable, less woody-textured taproot.

Lettuce

Lettuce

Lettuce is an annual plant of the family Asteraceae. It is most often grown as a leaf vegetable, but sometimes for its stem and seeds. Lettuce is most often used for salads, although it is also seen in other kinds of food, such as soups, sandwiches and wraps; it can also be grilled. One variety, celtuce, is grown for its stems, which are eaten either raw or cooked. In addition to its main use as a leafy green, it has also gathered religious and medicinal significance over centuries of human consumption. Europe and North America originally dominated the market for lettuce, but by the late 20th century the consumption of lettuce had spread throughout the world. As of 2017, world production of lettuce and chicory was 27 million tonnes, 56 percent of which came from China.

Bacteria

Bacteria

Bacteria are ubiquitous, mostly free-living organisms often consisting of one biological cell. They constitute a large domain of prokaryotic microorganisms. Typically a few micrometres in length, bacteria were among the first life forms to appear on Earth, and are present in most of its habitats. Bacteria inhabit soil, water, acidic hot springs, radioactive waste, and the deep biosphere of Earth's crust. Bacteria are vital in many stages of the nutrient cycle by recycling nutrients such as the fixation of nitrogen from the atmosphere. The nutrient cycle includes the decomposition of dead bodies; bacteria are responsible for the putrefaction stage in this process. In the biological communities surrounding hydrothermal vents and cold seeps, extremophile bacteria provide the nutrients needed to sustain life by converting dissolved compounds, such as hydrogen sulphide and methane, to energy. Bacteria also live in symbiotic and parasitic relationships with plants and animals. Most bacteria have not been characterised and there are many species that cannot be grown in the laboratory. The study of bacteria is known as bacteriology, a branch of microbiology.

Cellulose

Cellulose

Cellulose is an organic compound with the formula (C6H10O5)n, a polysaccharide consisting of a linear chain of several hundred to many thousands of β(1→4) linked D-glucose units. Cellulose is an important structural component of the primary cell wall of green plants, many forms of algae and the oomycetes. Some species of bacteria secrete it to form biofilms. Cellulose is the most abundant organic polymer on Earth. The cellulose content of cotton fiber is 90%, that of wood is 40–50%, and that of dried hemp is approximately 57%.

Song thrush

Song thrush

The song thrush is a thrush that breeds across the West Palearctic. It has brown upper-parts and black-spotted cream or buff underparts and has three recognised subspecies. Its distinctive song, which has repeated musical phrases, has frequently been referred to in poetry.

Frog

Frog

A frog is any member of a diverse and largely carnivorous group of short-bodied, tailless amphibians composing the order Anura. The oldest fossil "proto-frog" Triadobatrachus is known from the Early Triassic of Madagascar, but molecular clock dating suggests their split from other amphibians may extend further back to the Permian, 265 million years ago. Frogs are widely distributed, ranging from the tropics to subarctic regions, but the greatest concentration of species diversity is in tropical rainforest. Frogs account for around 88% of extant amphibian species. They are also one of the five most diverse vertebrate orders. Warty frog species tend to be called toads, but the distinction between frogs and toads is informal, not from taxonomy or evolutionary history.

Decollate snail

Decollate snail

The decollate snail, scientific name Rumina decollata, is a medium-sized predatory land snail, a species of terrestrial pulmonate gastropod mollusk in the family Achatinidae. It is a European species that has been introduced in a number of areas worldwide.

Polymita

Polymita

Polymita is a genus of large, air-breathing land snails, terrestrial pulmonate gastropod mollusks in the family Helminthoglyptidae.

Hawaii

Hawaii

Hawaii is a state in the Western United States, about 2,000 miles (3,200 km) from the U.S. mainland in the Pacific Ocean. It is the only U.S. state outside North America, the only state that is an archipelago, and the only state in the tropics.

Edible snails

H. pomatia and H. aspersa are the two edible species that are most used in European cuisine. Spanish cuisine also uses Otala punctata, Theba pisana, and Iberus gualterianus alonensis, amongst others. The process of snail farming is called heliciculture.

Escargots are often traditionally served as appetizers. They may also be used as ingredients in other recipes.

Snails contain many nutrients. They are rich in calcium and also contain vitamin B1 and E. They contain various essential amino acids, and are low in calories and fat.

Discover more about Edible snails related topics

Cuisine

Cuisine

A cuisine is a style of cooking characterized by distinctive ingredients, techniques and dishes, and usually associated with a specific culture or geographic region. Regional food preparation techniques, customs, and ingredients combine to enable dishes unique to a region.

Otala punctata

Otala punctata

Otala punctata is a species of air-breathing land snail, a terrestrial pulmonate gastropod mollusk in the family Helicidae.

Theba pisana

Theba pisana

Theba pisana, common names the white garden snail, sand hill snail, white Italian snail, Mediterranean coastal snail, and simply just the Mediterranean snail, is an edible species of medium-sized, air-breathing land snail, a terrestrial pulmonate gastropod mollusk in the family Helicidae, the typical snails.

Heliciculture

Heliciculture

Heliciculture, commonly known as snail farming, is the process of raising edible land snails, primarily for human consumption or cosmetic use. The meat and snail eggs a.k.a. white caviar can be consumed as escargot and as a type of caviar, respectively.

Escargot

Escargot

Snails are considered edible in certain areas such as the Mediterranean region, Africa, or Southeast Asia, while in other cultures it is considered as taboo food. In American English, edible land snails are also called escargot, taken from the French word for 'snail', and the production of snails for consumption is called snail farming or heliciculture. Snails as food date back to ancient times, and it was a dish that was already served on the tables of Ancient Greece and Rome.

Calcium

Calcium

Calcium is a chemical element with the symbol Ca and atomic number 20. As an alkaline earth metal, calcium is a reactive metal that forms a dark oxide-nitride layer when exposed to air. Its physical and chemical properties are most similar to its heavier homologues strontium and barium. It is the fifth most abundant element in Earth's crust, and the third most abundant metal, after iron and aluminium. The most common calcium compound on Earth is calcium carbonate, found in limestone and the fossilised remnants of early sea life; gypsum, anhydrite, fluorite, and apatite are also sources of calcium. The name derives from Latin calx "lime", which was obtained from heating limestone.

Vitamin E

Vitamin E

Vitamin E is a group of eight fat soluble compounds that include four tocopherols and four tocotrienols. Vitamin E deficiency, which is rare and usually due to an underlying problem with digesting dietary fat rather than from a diet low in vitamin E, can cause nerve problems. Vitamin E is a fat-soluble antioxidant which may help protect cell membranes from reactive oxygen species. Worldwide, government organizations recommend adults consume in the range of 3 to 15 mg per day. As of 2016, consumption was below recommendations according to a worldwide summary of more than one hundred studies that reported a median dietary intake of 6.2 mg per day for alpha-tocopherol.

Amino acid

Amino acid

Amino acids are organic compounds that contain both amino and carboxylic acid functional groups. Although over 500 amino acids exist in nature, by far the most important are the alpha-amino acids, which comprise proteins. Only 22 alpha amino acids appear in the genetic code.

Calorie

Calorie

The calorie is a unit of energy that originated from the obsolete caloric theory of heat. For historical reasons, two main definitions of "calorie" are in wide use. The large calorie, food calorie, dietary calorie, or kilogram calorie was originally defined as the amount of heat needed to raise the temperature of one kilogram of water by one degree Celsius. The small calorie or gram calorie was defined as the amount of heat needed to cause the same increase in one gram of water. Thus, 1 large calorie is equal to 1000 small calories.

Fat

Fat

In nutrition, biology, and chemistry, fat usually means any ester of fatty acids, or a mixture of such compounds, most commonly those that occur in living beings or in food.

List of Helix species

Scientific name[a] IUCN Red List Status Distribution Picture
Helix albescens
Rossmässler, 1839
LC IUCN Helix-albescens-map-eur-nm-moll.jpg 2015-07-02-0668(0) (19341856612).jpg
Helix anctostoma
Martens, 1874
Helix antiochiensis
Kobelt, 1896
Helix tripolitana (MNHN-IM-2000-27767).jpeg
Helix aporina
Castro, 1887
Helix aporina (MNHN-IM-2000-30109).jpeg
Helix asemnis
Bourguignat, 1860
LC IUCN Helix-asemnis-map-eur-nm-moll.jpg
Helix borealis
Mousson, 1859
DD IUCN
Helix buchii
(Dubois de Montpéreux, 1840)
Helix calabrica
Westerlund, 1876
Helix ceratina
Shuttleworth, 1843
Tyrrhenaria-ceratina-map-eur-nm-moll.jpg Tyrrhenaria ceratina (MNHN-IM-2010-13180).jpeg
Helix cincta
O. F. Müller, 1774
LC IUCN Helix-cincta-map-eur-nm-moll.jpg Helix cincta cincta 01.JPG
Helix dormitoris
Kobelt, 1898
Helix-dormitoris-map-eur-nm-moll.jpg
Helix engaddensis
Bourguignat, 1852
Helix engaddensis 2019.jpg
Helix escherichi
O. Boettger, 1898
Helix-escherichi-map-eur-nm-moll.jpg
Helix fathallae
Nägele, 1901
Helix figulina
Rossmässler, 1839
LC IUCN Helix-figulina-map-eur-nm-moll.jpg 1k Helix-figulina 01.jpg
Helix gussoneana
L. Pfeiffer, 1848
Helix hedenborgi
L. Pfeiffer, 1846
Helix kazouiniana
Pallary, 1939
Helix krejcii
Wenz in Krejci-Graf & Wenz, 1926
Helix ligata
O. F. Müller, 1774
DD IUCN Helix-ligata-map-eur-nm-moll.jpg
Helix lucorum
Linnaeus, 1758
Helix-lucorum-map-eur-nm-moll.jpg Helix lucorum 2.jpg
Helix lutescens
Rossmässler, 1837
LC IUCN Helix-lutescens-map-eur-nm-moll.jpg Slimak lutescens.jpg
Helix melanostoma
Draparnaud, 1801
Helix-melanostoma-map-eur-nm-moll.jpg Helix melanostoma 01.JPG
Helix mileti
Kobelt, 1906
Helix-mileti-map-eur-nm-moll.jpg
Helix mrazeci
Sevastos, 1922
Helix nicaeensis
A. Férussac, 1821
Helix nucula
Mousson, 1854
LC IUCN Helix-nucula-map-eur-nm-moll.jpg Helix nucula shell(s) 06.jpg
Helix pachya
Bourguignat, 1860
Helix pathetica
Mousson, 1854
Helix pelagonesica
(Rolle, 1898)
Helix philibinensis
Rossmässler, 1839
LC IUCN Helix philibinensis 01.JPG
Helix pomacella
Mousson, 1854
LC IUCN Helix-pomacella-map-eur-nm-moll.jpg
Helix pomatella
Kobelt, 1876
Helix pomatia
Linnaeus, 1758
Helix pomatia 89a.jpg
Helix pronuba
Westerlund & Blanc, 1879
Helix salomonica
Nägele, 1899
Helix schlaeflii
Mousson, 1859
Helix secernenda
Rossmässler, 1847
LC IUCN Helix-secernenda-map-eur-nm-moll.jpg
Helix straminea
Briganti, 1825
LC IUCN
Helix sublutescens
Wenz in Krejci & Wenz, 1926
Helix thessalica
O. Boettger, 1886
LC IUCN
Helix tourannensis
Souleyet, 1852
Helix valentini
Kobelt, 1891
EN IUCN Helix-valentini-map-eur-nm-moll.jpg
Helix vladica
(Kobelt, 1898)
LC IUCN

There are also 6 extinct species of Helix:

Discover more about List of Helix species related topics

Helix albescens

Helix albescens

Helix albescens is a species of gastropods belonging to the family Helicidae.

Helix asemnis

Helix asemnis

Helix asemnis is a species of gastropods belonging to the family Helicidae.

Helix ceratina

Helix ceratina

Helix ceratina is a species of air-breathing land snail, a terrestrial pulmonate gastropod mollusk in the family Helicidae, the typical snails.

Helix cincta

Helix cincta

Helix cincta is a species of gastropods belonging to the family Helicidae.

Helix engaddensis

Helix engaddensis

Helix engaddensis is a species of snail common in the Levant, both in Mediterranean, desert and montane climates. It is smaller than the closely related European Garden snail and usually lighter in color. H. engaddensis goes through estivation. It is dormant in the ground during the dry season and emerges after the first rains. Mating takes place soon after emerging. The snails are active through winter and return to an inactive state at the end of the wet season (midspring).

Helix figulina

Helix figulina

Helix figulina is a species of air-breathing land snail, a terrestrial pulmonate gastropod mollusk in the family Helicidae, the typical snails.

Helix lucorum

Helix lucorum

Helix lucorum is a species of large, edible, air-breathing land snail, a terrestrial pulmonate gastropod mollusk in the family Helicidae, the typical snails.

Helix lutescens

Helix lutescens

Helix lutescens is species of air-breathing land snail, a terrestrial pulmonate gastropod mollusk in the family Helicidae, the true snails.

Helix melanostoma

Helix melanostoma

Helix melanostoma is a species of air-breathing land snail, a terrestrial pulmonate gastropod mollusk in the subfamily Helicinae of the family Helicidae, the typical snails.

Helix nucula

Helix nucula

Helix nucula is a species of air-breathing land snail, a terrestrial pulmonate gastropod mollusk in the family Helicidae, the typical snails. According to the malacologist E. Neubert, the taxonomy of this species is rather complex; the species has been confused with Helix figulina, a related species. Scientists who conducted a study using mitochondrial DNA sequences confirmed that H. nucula is a distinct monophyletic species, and within that clade are two lineages that were well distinguished, one from the North African population and the other from southwest Crete.

Helix pomatia

Helix pomatia

Helix pomatia, common names the Roman snail, Burgundy snail, or escargot, is a species of large, edible, air-breathing land snail, a pulmonate gastropod terrestrial mollusc in the family Helicidae. It is one of Europe's biggest species of land snail.

Taxon inquirendum

A long list
  • Helix (Cochlichondra) Jan, 1830
  • Helix (Cochlogibba) Jan, 1830
  • Helix (Patera) Albers, 1850 (Invalid: junior homonym of Patera Lesson, 1839 [Cnidaria])
  • Helix achilli Bourguignat, 1883
  • Helix acompsia (Bourguignat, 1863)
  • Helix adelaidae L. Pfeiffer, 1857
  • Helix adusta Hinds, 1845
  • Helix ahmedi Pallary, 1899
  • Helix aidae Pallary, 1899
  • Helix alata L. Pfeiffer, 1856
  • Helix albersi L. Pfeiffer, 1849
  • Helix albidens Benson, 1853
  • Helix alinae Pallary, 1899
  • Helix alsiella Pallary, 1899
  • Helix alveolus Heude, 1890
  • Helix annamitica Crosse & Fischer, 1863
  • Helix anomia Heude, 1890
  • Helix ansorinus Theobald, 1866
  • Helix appressa Say, 1821
  • Helix arabophila Pallary, 1898
  • Helix araneaetela Heude, 1885
  • Helix artificiosa Benson, 1856
  • Helix arundinetorum Heude, 1882
  • Helix avidorum Cox, 1868
  • Helix bahamensis L. Pfeiffer, 1845
  • Helix barclayana L. Pfeiffer, 1847
  • Helix barrakporensis Benson, 1859
  • Helix batchianensis L. Pfeiffer, 1861
  • Helix batesii L. Pfeiffer, 1860
  • Helix beccarii Jickeli, 1874 (taxon inquirendum, replacement name for Helix ciliata non Venetz)
  • Helix belcheri L. Pfeiffer, 1846
  • Helix bicallosula Heude, 1886
  • Helix biconvexa Martens, 1864
  • Helix binneyana L. Pfeiffer, 1847
  • Helix bizona Gredler, 1884
  • Helix boissieri S. Moricand, 1846
  • Helix bombax Benson, 1859
  • Helix bonplandii Lamarck, 1822
  • Helix bourguignati L. Pfeiffer, 1857
  • Helix boxalli G. B. Sowerby III, 1888 (use in recent literature currently undocumented)
  • Helix brocchii Jickeli, 1874 (invalid; not F. Sandberger, 1872)
  • Helix brucei Jickeli, 1874 (use in recent literature currently undocumented)
  • Helix bullina Férussac, 1832 (has also been mistakenly identified as Amphibulima rubescens Deshayes, 1830) [15]
  • Helix buxina Heude, 1886
  • Helix caelatura Férussac, 1821
  • Helix calpis Benson, 1859
  • Helix calymma Schmacker & O. Boettger, 1894
  • Helix campelica Mabille, 1886 (taxon inquirendum, no documented use in recent literature)
  • Helix capessens Benson, 1856
  • Helix carinata Link, 1807
  • Helix carmeliensis L. Pfeiffer, 1861
  • Helix carpalima J. Mabille, 1889
  • Helix cassidula Benson, 1859
  • Helix caucasica L. Pfeiffer, 1846
  • Helix ceryx Benson, 1859
  • Helix chadiana Pallary, 1899
  • Helix cherifiana Pallary, 1899
  • Helix chionodiscus L. Pfeiffer, 1857
  • Helix cimex Reeve, 1854
  • Helix coagulata L. Pfeiffer, 1856
  • Helix conrauxiana Heude, 1885
  • Helix conulus Martens, 1864
  • Helix corneola Clessin, 1874
  • Helix cumulus Reeve, 1854
  • Helix cyclostomopsis I. Lea, 1834
  • Helix cyclotrema Benson, 1863
  • Helix cypsele L. Pfeiffer, 1849
  • Helix cyrenaica E. von Martens, 1883
  • Helix debauxii Noulet, 1854
  • Helix decidua L. Pfeiffer, 1857
  • Helix delavayana Heude, 1885
  • Helix demolita Heude, 1885
  • Helix depsta Cox, 1870
  • Helix dhericourtiana Bourguignat, 1885
  • Helix diaphana I. Lea, 1834
  • Helix diniensis Rambur, 1868
  • Helix doufourii Grateloup, 1840
  • Helix dubia Clessin, 1874
  • Helix ductilis Pfeiffer, 1857
  • Helix egregia Deshayes, 1850
  • Helix emmae Pallary, 1899
  • Helix exacta L. Pfeiffer, 1863
  • Helix exilis O. F. Müller, 1774
  • Helix expansa Clessin, 1874
  • Helix exserta L. Pfeiffer, 1856
  • Helix faradensis L. Pfeiffer, 1861
  • Helix fauna Philippi, 1851
  • Helix faunus Broderip, 1841
  • Helix ferretiana Bourguignat, 1883
  • Helix filaris L. Pfeiffer, 1845
  • Helix flammata Férussac, 1821
  • Helix flaveola Martens, 1864
  • Helix fornicata Gould, 1846
  • Helix forrestiana Angas, 1875
  • Helix fortunei Heude, 1882
  • Helix fossaria Montagu, 1803
  • Helix fritillata Benson, 1863
  • Helix galinieriana Bourguignat, 1883
  • Helix gaudiella Mabille, 1886 (taxon inquirendum, no documented use in recent literature)
  • Helix gerlachi Möllendorff
  • Helix gigas L. Pfeiffer, 1850
  • Helix giraudeliana Heude, 1882
  • Helix globula I. Lea, 1834
  • Helix glomerosa Godwin-Austen, 1883
  • Helix gordoniae Benson, 1863
  • Helix graminum Heude, 1882
  • Helix griffithi L. Pfeiffer, 1846
  • Helix grossularia L. Pfeiffer, 1861
  • Helix guadalcanarensis Cox, 1872
  • Helix guinaria L. Pfeiffer, 1861
  • Helix haesitans Heude, 1888
  • Helix hamacenica Bourguignat, 1883
  • Helix hamudae Kobelt, 1886 (taxon inquirendum, use in recent literature currently undocumented)
  • Helix helmii G. B. Sowerby I, 1838 (taxon inquirendum, use in recent literature currently undocumented)
  • Helix herbini Bourguignat, 1883
  • Helix herpestes Heude, 1885
  • Helix herza J. Mabille, 1889
  • Helix hians L. Pfeiffer, 1846
  • Helix hongkongensis Deshayes, 1874
  • Helix horripilosella Heude, 1885
  • Helix huberiana Heude, 1882
  • Helix iae Pallary, 1899
  • Helix improvisa Heude, 1885
  • Helix ingloria Heude, 1890
  • Helix involuta L. Pfeiffer, 1845
  • Helix jacob Gredler, 1887
  • Helix keratina Heude, 1889
  • Helix kuangtunensis Gredler, 1881
  • Helix labiata L. Pfeiffer, 1845
  • Helix lamellata M. Bielz, 1851
  • Helix languescens Heude, 1890
  • Helix lariollei Pallary, 1899
  • Helix latrunculorum Heude, 1885
  • Helix lenticularis Morelet, 1853
  • Helix linnaeana L. Pfeiffer, 1845
  • Helix listeri Broderip, 1841
  • Helix loxodon L. Pfeiffer, 1850
  • Helix luctativa J. Mabille, 1889
  • Helix lutuosa Deshayes, 1874
  • Helix lyrata Gould, 1846
  • Helix madritensis Rambur, 1868
  • Helix mamilla
  • Helix marcida Benson, 1853
  • Helix margaritacea A. Schmidt, 1852
  • Helix melillensis Pallary, 1899
  • Helix mellita Heude, 1886
  • Helix mellitula Heude, 1886
  • Helix mendanae Cox, 1873
  • Helix mendicaria L. Pfeiffer, 1860
  • Helix mensalis Heude, 1888
  • Helix meretrix G. B. Sowerby I, 1841 (taxon inquirendum, use in recent literature currently undocumented)
  • Helix meridionalis Mousson, 1854 (taxon inquirendum, non Wood, 1828)
  • Helix mersispira Martens, 1864
  • Helix mesquiniana Pallary, 1898
  • Helix micacea Heude, 1882
  • Helix microzaffarina Pallary, 1898
  • Helix mighelsiana L. Pfeiffer, 1847
  • Helix miliaria Gredler, 1881
  • Helix mimicula Heude, 1888
  • Helix minutella J. Mabille, 1889
  • Helix monodonta I. Lea, 1834
  • Helix moquiniana Raymond, 1853
  • Helix moretonensis Reeve, 1854
  • Helix muscarum I. Lea, 1834
  • Helix mystagoga J. Mabille, 1888
  • Helix nautarum Heude, 1882
  • Helix nicolai Klećak, 1880 (taxon inquirendum, use in more recent taxonomic literature currently undocumented)
  • Helix novare L. Pfeiffer, 1861
  • Helix nystiana L. Pfeiffer, 1846
  • Helix oberndoerferi Kobelt, 1882 (taxon inquirendum, use in more recent taxonomic literature currently undocumented)
  • Helix obscura Deshayes, 1874
  • Helix oleosa L. Pfeiffer, 1850
  • Helix omissa L. Pfeiffer, 1856
  • Helix onisterella J. Mabille, 1889
  • Helix opaca Gmelin, 1791
  • Helix ophelia Reeve, 1854
  • Helix ovumreguli I. Lea, 1834
  • Helix pansa Benson, 1856
  • Helix pantheia J. Mabille, 1888
  • Helix papillata L. Pfeiffer, 1846
  • Helix papillionacea Valenciennes, 1827
  • Helix papyracea Broderip, 1841
  • Helix paraeruginosa Heude, 1888
  • Helix parasitarum Heude, 1885
  • Helix parasitica Heude, 1885
  • Helix paropta Mabille, 1886 (taxon inquirendum, no documented use in recent literature)
  • Helix paulinae Pallary, 1899
  • Helix peguensis Benson, 1860
  • Helix pemphigodes L. Pfeiffer, 1847
  • Helix pena Cox, 1868
  • Helix penicillata Gould, 1842
  • Helix pennantiana L. Pfeiffer, 1845
  • Helix perforata Deshayes, 1874
  • Helix permellita Heude, 1890
  • Helix phragmitum Heude, 1882
  • Helix phyllophaga Heude, 1882
  • Helix pilidion Benson, 1860
  • Helix pilifera E. von Martens, 1869
  • Helix piligera Gredler, 1885
  • Helix plicatilis Deshayes, 1870
  • Helix pliculosa L. Pfeiffer, 1857
  • Helix portoricensis L. Pfeiffer, 1847
  • Helix praelongata Pallary, 1898
  • Helix praeruginosa Heude, 1888
  • Helix probata Mabille, 1898
  • Helix prshewalskii Martens, 1881
  • Helix prunum Férussac, 1821
  • Helix puberosula Heude, 1885
  • Helix purpuragula I. Lea, 1834
  • Helix puteolus Benson, 1853
  • Helix pyantha J. Mabille, 1889
  • Helix pyramidalis G. B. Sowerby I, 1841 (taxon inquirendum, use in recent literature currently undocumented)
  • Helix pyxis Hinds, 1845
  • Helix quedenfeldti E. von Martens, 1890
  • Helix quirosi Cox, 1873
  • Helix radicicola Benson, 1859
  • Helix radulella Heude, 1885
  • Helix radulina Heude, 1888
  • Helix rebellis Heude, 1885
  • Helix reginae W. J. Broderip, 1841
  • Helix renaltiana Heude, 1889
  • Helix rerayana Pallary, 1899
  • Helix rissoana L. Pfeiffer, 1847
  • Helix rosacea G. B. Sowerby I, 1839 (taxon inquirendum, Invalid: junior homonym of Helix rosacea O. F. Müller, 1774)
  • Helix rostrata L. Pfeiffer, 1847
  • Helix rupelli Deshayes, 1870
  • Helix samara Heude, 1886
  • Helix samarella Heude, 1888
  • Helix sanata Heude, 1889
  • Helix sansitus Cox, 1870
  • Helix saturnia Gould, 1846
  • Helix savignyana Ehrenberg, 1831
  • Helix saxatilis Gould, 1846 (taxon inquirendum, not Hartmann, 1821)
  • Helix scenoma Benson, 1863
  • Helix schweinfurthi E. von Martens, 1877
  • Helix secundaria Heude, 1890
  • Helix sedentaria Heude, 1885
  • Helix seguiniana Heude, 1885
  • Helix semicarinata Ancey, 1881
  • Helix semifusca Deshayes, 1832
  • Helix semigranosa G. B. Sowerby I, 1841 (taxon inquirendum, use in recent literature currently undocumented)
  • Helix semirugosa Pallary, 1899
  • Helix sempriniana Heude, 1882
  • Helix sireti Pallary, 1898
  • Helix sororia Cox, 1870
  • Helix specialis Bourguignat, 1881
  • Helix spengleriana L. Pfeiffer, 1847
  • Helix spirillus Gould, 1852
  • Helix squalus Hinds, 1845
  • Helix squamosella Heude, 1882
  • Helix squamulosa Mousson, 1856
  • Helix stauropolitana A. Schmidt, 1855
  • Helix strucki Maltzan, 1886 (taxon inquirendum, use in recent literature currently undocumented)
  • Helix subcinctula Heude, 1890
  • Helix subechinata Deshayes, 1870
  • Helix subgranosa Le Guillou, 1842
  • Helix subgriseola Heude, 1888
  • Helix sublallementiana Pallary, 1899
  • Helix subnivellina Bourguignat, 1883
  • Helix subnuda J. Mabille, 1889
  • Helix subparasitica Heude, 1888
  • Helix suffulta Benson, 1853
  • Helix sumatrana Martens, 1864
  • Helix supracostata Kobelt, 1882 (taxon inquirendum, use in recent literature currently undocumented)
  • Helix taliensis Heude, 1890
  • Helix talifouensis Heude, 1888
  • Helix tarifensis Bourguignat in Servain, 1881
  • Helix tarnieri Pallary, 1899
  • Helix tenuis L. Pfeiffer, 1845
  • Helix terrestris Forskål, 1775
  • Helix tescorum Benson, 1853
  • Helix theobryta J. Mabille, 1889
  • Helix thibetica Deshayes, 1870
  • Helix tickelli Theobald, 1859
  • Helix tristrami L. Pfeiffer, 1860
  • Helix tumida L. Pfeiffer, 1846
  • Helix turbinella Heude, 1890
  • Helix tutuillae Cox, 1870
  • Helix vannaelaeve Cox, 1870
  • Helix variolosa L. Pfeiffer, 1846
  • Helix vatheleti J. Mabille, 1888
  • Helix vesica I. Lea, 1834
  • Helix vesica L. Pfeiffer, 1842 (taxon inquirendum, not Helix vesica I. Lea, 1834)
  • Helix vicinella Heude, 1890
  • Helix vitiensis L. Pfeiffer, 1855
  • Helix vitreola Heude, 1890
  • Helix vivacula J. Mabille, 1889
  • Helix vorticellina Heude, 1889
  • Helix wanganensis Cox, 1870
  • Helix welschi Pallary, 1899
  • Helix woodiana I. Lea, 1834
  • Helix yentaiensis Crosse & Debeaux
  • Helix yorkensis Reeve, 1854
  • Helix zelina Cox, 1873
  • Helix zeus L. Pfeiffer, 1843
  • Helix zollingeri L. Pfeiffer, 1854
  • Helix zonites L. Pfeiffer, 1846
  • Helix zorica J. Mabille, 1889
  • Helix zoroaster Theobald, 1859

Nomen dubium

  • Helix calomorpha Jonas, 1839
  • Helix cornea Draparnaud, 1801
  • Helix laevigata Linnaeus, 1767 accepted as Velutina laevigata (Linnaeus, 1767) (nomen dubium, original combination)
  • Helix minutialis Deshayes, 1851
  • Helix multispirata Hombron & Jacquinot, 1852
  • Helix oceanica Le Guillou, 1842
  • Helix rotula Hombron & Jacquinot, 1852 (nomen dubium, invalid: junior homonym of Helix rotula Lowe, 1831; no replacement name available)
  • Helix rugosa L. Pfeiffer, 1843
  • Helix undulata Gmelin, 1791
  • Helix unispiralis Montagu, 1803

Nomen nudum

Subgenus Helix (Pitys) H. Beck, 1837

  • Helix (Pitys) oparana H. Beck, 1837
  • Helix angulata Férussac, 1821
  • Helix congellana Krauss in E. von Martens, 1860
  • Helix exoptata Tate, 1882
  • Helix globula Krynicki, 1838
  • Helix ingrami Blanford, 1876
  • Helix roseri F. Krauss in E. von Martens, 1860
  • Helix roseri Krauss in E. von Martens, 1860
  • Helix somersetii Prime, 1853
  • Helix tingitana Beck, 1837
  1. ^ A binomial authority in parentheses indicates that the species was originally described in a genus other than Helix .

Source: "Helix (gastropod)", Wikipedia, Wikimedia Foundation, (2022, December 9th), https://en.wikipedia.org/wiki/Helix_(gastropod).

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References
  1. ^ a b Rosenberg, G.; Bouchet, P. (2014). Helix Linnaeus, 1758. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=153970 on 2015-02-24
  2. ^ Ivanov M., Hrdličková, S. & Gregorová, R. (2001) Encyklopedie zkamenělin. – Rebo Productions, Dobřejovice, 1. vydání, 312 pp., p. 126. (in Czech)
  3. ^ A. Eastham, Alastair Small, Michael Ross MadceqrefvrevrecKinnon, Stephen G. Monckton, David S. Reese, Robert J. Buck (2002) The Excavations of San Giovanni Di Ruoti: The Faunal and Plant Remains, University of Toronto Press, 232 pages ISBN 0-8020-4865-X
  4. ^ Helix pomatia Linnaeus, 1758 in GBIF Secretariat (2017). GBIF Backbone Taxonomy. Checklist dataset https://doi.org/10.15468/39omei accessed via GBIF.org on 2018-12-08.
  5. ^ Rumi, A., Sánchez, J., & Ferrando, N. S. (2010). Theba pisana (Müller, 1774) (Gastropoda, Helicidae) and other alien land molluscs species in Argentina. Biological Invasions, 12(9), 2985–2990. doi:10.1007/s10530-010-9715-x
  6. ^ USDA (Robinson D.G., Redmond L. and Hennessey R.) (1998). Importation and interstate movement of live, edible land snails: Cantareus apertus (Born), Cryptomphalus aspersus (Müller), Eobania vermiculata (Müller), Helix pomatia Linné, and Otala lactea (Müller) (Pulmonata: Helicidae): qualitative pest risk assessment.
  7. ^ Herbert D.G. 2010. The introduced terrestrial mollusca of South Africa. Published by South African National Biodiversity Institute Libraries. Vol. 15. pp. 120.
  8. ^ The Cornu Problem Archived 27 September 2007 at the Wayback Machine
  9. ^ Westerlund, C. A. 1889. Fauna der in der paläarctischen Region (Europa, Kaukasien, Sibirien, Turan, Persien, Kurdistan, Armenien, Mesopotamien, Kleinasien, Syrien, Arabien, Egypten, Tripolis, Tunesien, Algerien und Marocco) lebenden Binnenconchylien. II. Gen. Helix. - pp. 1-473, 1-31, 1-8. Berlin. (Friedländer).
  10. ^ Pfeiffer, L. & Clessin, S. 1881. Nomenclator heliceorum viventium quo continetur nomina omnium hujus familiae generum et specierum hodie cognitarum, disposita ex affinitate naturali. - pp. 1-617. Cassellis. (Fischer).
  11. ^ The Roman Snail, Helix pomatia Archived 24 November 2005 at the Wayback Machine
  12. ^ Willoughby, David P. (1974). "Running and Jumping". Natural History. 83 (3): 71.
  13. ^ Yee, Angie (1999). "Speed of a Snail". The Physics Factbook. Archived from the original on 4 February 2012.
  14. ^ Cameron, Robert (2016). Slugs and Snails. Collins New Naturalist Library, Book 133. HarperCollins.
  15. ^ Nathaniel F. Shoobs | Bram Breure, Revision-of-the-Lesser-Antillean-genera-Amphibulima-and-Pellicula
    • Helix buvignieri Deshayes, 1874]

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