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Heliciculture

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A snail farm near Eyragues, Provence, France
A snail farm near Eyragues, Provence, France

Heliciculture, commonly known as snail farming, is the process of raising edible land snails, primarily for human consumption or cosmetic use.[1] The meat and snail eggs a.k.a. white caviar can be consumed as escargot and as a type of caviar, respectively.[2]

Perhaps the best-known edible land snail species in the Western world is Helix pomatia, commonly known as the Roman snail or the Burgundy snail. This species, however, is not fit for profitable snail farming, and is normally harvested from nature.

Commercial snail farming in the Western world is typically done with Cornu aspersum (morphotypically divided into C. a. aspersa and C. a. maxima), formerly known as Helix aspersa. In tropical climates, snail farming is typically done with the African snail, which biologically does not fall in the family Helicidae, so the meat of which may not be called escargot.

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Land snail

Land snail

A land snail is any of the numerous species of snail that live on land, as opposed to the sea snails and freshwater snails. Land snail is the common name for terrestrial gastropod mollusks that have shells. However, it is not always easy to say which species are terrestrial, because some are more or less amphibious between land and fresh water, and others are relatively amphibious between land and salt water.

Snail caviar

Snail caviar

Snail caviar, also known as escargot caviar or escargot pearls, is a type of caviar that consists of fresh or processed eggs of land snails. It is a luxury gourmet speciality produced in France and Poland. They were also a delicacy in the ancient world, also known as "Pearls of Aphrodite" for their supposed aphrodisiac properties.

Escargot

Escargot

Snails are considered edible in certain areas such as the Mediterranean region, Africa, or Southeast Asia, while in other cultures it is considered as taboo food. In American English, edible land snails are also called escargot, taken from the French word for 'snail', and the production of snails for consumption is called snail farming or heliciculture. Snails as food date back to ancient times, and it was a dish that was already served on the tables of Ancient Greece and Rome.

Western world

Western world

The Western world, also known as the West, primarily refers to various nations and states in the regions of Australasia, Europe, and the Americas. The Western world likewise is called the Occident, in contrast to the Eastern world known as the Orient. The West is considered an evolving concept; made up of cultural, political, and economic synergy among diverse groups of people, and not a rigid region with fixed borders and members. Definitions for "Western world" vary according to context and perspectives.

Helix pomatia

Helix pomatia

Helix pomatia, common names the Roman snail, Burgundy snail, or escargot, is a species of large, edible, air-breathing land snail, a pulmonate gastropod terrestrial mollusc in the family Helicidae. It is one of Europe's biggest species of land snail.

Cornu aspersum

Cornu aspersum

Cornu aspersum, known by the common name garden snail, is a species of land snail in the family Helicidae, which includes some of the most familiar land snails. Of all terrestrial molluscs, this species may well be the most widely known. It was classified under the name Helix aspersa for over two centuries, but the prevailing classification now places it in the genus Cornu.

Helicidae

Helicidae

Helicidae is a large, diverse family of small to large, air-breathing land snails, sometimes called the "typical snails."

History

Roasted snail shells have been found in archaeological excavations, an indication that snails have been eaten since prehistoric times.[3][4]

Lumaca romana, (translation: Roman snail), was an ancient method of snail farming or heliciculture in the region about Tarquinia. This snail-farming method was described by Fulvius Lippinus (49 BC) and mentioned by Marcus Terentius Varro in De Re rustica III, 12. The snails were fattened for human consumption using spelt and aromatic herbs. People usually raised snails in pens near their houses, and these pens were called "cochlea".[5]

The Romans, in particular, are known to have considered escargot as an elite food, as noted in the writings of Pliny the Elder. The Romans selected the best snails for breeding. Fulvius Lippinus started this practice. Various species were consumed by the Romans. Shells of the edible land snail species Otala lactea have been recovered in archaeological excavations of Volubilis in present-day Morocco.[6]

"Wallfish" were also often eaten in Britain, but were never as popular as on the continent. There, people often ate snails during Lent, and in a few places, they consumed large quantities of snails at Mardi Gras or Carnival, prior to Lent.

According to some sources, the French exported brown garden snails to California in the 1850s, raising them as the delicacy escargot. Other sources claim that Italian immigrants were the first to bring the snail to the United States.[7]

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Tarquinia

Tarquinia

Tarquinia, formerly Corneto, is an old city in the province of Viterbo, Lazio, Central Italy, known chiefly for its ancient Etruscan tombs in the widespread necropoleis, or cemeteries, for which it was awarded UNESCO World Heritage status.

Marcus Terentius Varro

Marcus Terentius Varro

Marcus Terentius Varro was a Roman polymath and a prolific author. He is regarded as ancient Rome's greatest scholar, and was described by Petrarch as "the third great light of Rome". He is sometimes called Varro Reatinus to distinguish him from his younger contemporary Varro Atacinus.

Spelt

Spelt

Spelt, also known as dinkel wheat or hulled wheat, is a species of wheat that has been cultivated since approximately 5000 BC.

Pliny the Elder

Pliny the Elder

Gaius Plinius Secundus, called Pliny the Elder, was a Roman author, naturalist and natural philosopher, and naval and army commander of the early Roman Empire, and a friend of the emperor Vespasian. He wrote the encyclopedic Naturalis Historia, which became an editorial model for encyclopedias. He spent most of his spare time studying, writing, and investigating natural and geographic phenomena in the field.

Quintus Fulvius Lippinus

Quintus Fulvius Lippinus

Quintus Fulvius Lippinus, Fulvius Lippinus for short was an enterprising Roman farmer from the first century BC. He lived in the Roman region of Tarquinia, today's Italian Tuscany. His dealings are described in the Rerum rusticarum libri III by Marcus Terentius Varro, and a century later in Pliny the Elder's Naturalis Historia.

Otala lactea

Otala lactea

Otala lactea, known as the milk snail or Spanish snail, is a large, edible species of air-breathing land snail, a terrestrial pulmonate gastropod mollusk, in the family Helicidae, the typical snails.

Volubilis

Volubilis

Volubilis is a partly-excavated Berber-Roman city in Morocco situated near the city of Meknes that may have been the capital of the Kingdom of Mauretania, at least from the time of King Juba II. Before Volubilis, the capital of the kingdom may have been at Gilda.

Morocco

Morocco

Morocco, officially the Kingdom of Morocco, is a country in the Maghreb region of North Africa. It overlooks the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria to the east, and the disputed territory of Western Sahara to the south. Mauritania lies to the south of Western Sahara. Morocco also claims the Spanish exclaves of Ceuta, Melilla and Peñón de Vélez de la Gomera, and several small Spanish-controlled islands off its coast. It spans an area of 446,300 km2 (172,300 sq mi) or 710,850 km2 (274,460 sq mi), with a population of roughly 37 million. Its official and predominant religion is Islam, and the official languages are Arabic and Berber; the Moroccan dialect of Arabic and French are also widely spoken. Moroccan identity and culture is a mix of Arab, Berber, and European cultures. Its capital is Rabat, while its largest city is Casablanca.

Lent

Lent

Lent is the solemn Christian religious observance in the liturgical year commemorating the 40 days Jesus Christ spent fasting in the desert and enduring temptation by Satan, according to the Gospels of Matthew, Mark and Luke, before beginning his public ministry. Lent is observed in the Anglican, Eastern Orthodox, Lutheran, Methodist, Moravian, Oriental Orthodox, Church of the East, United Protestant and Roman Catholic traditions of Christianity. Some Anabaptist, Baptist, Reformed, and nondenominational Christian churches also observe Lent, although many churches in these traditions do not.

Escargot

Escargot

Snails are considered edible in certain areas such as the Mediterranean region, Africa, or Southeast Asia, while in other cultures it is considered as taboo food. In American English, edible land snails are also called escargot, taken from the French word for 'snail', and the production of snails for consumption is called snail farming or heliciculture. Snails as food date back to ancient times, and it was a dish that was already served on the tables of Ancient Greece and Rome.

Edible land snail species

Three different species of snails for sale in a market in Turin, Italy
Three different species of snails for sale in a market in Turin, Italy

Most land snails are edible provided they are properly cooked. Their flavour varies by species and the way/method of cooking, and preferences may vary by culture. Only a few species are suitable for profitable farming.[8]

Edible land snails range in size from about 2 millimetres (564 in) long to the giant African snails, which occasionally grow up to 312 mm (1 ft 14 in) in length. "Escargot" most commonly refers to either Cornu aspersum or to Helix pomatia, although other varieties of snails are eaten. Lissachatina fulica, a giant African snail, is sliced and canned and passed off on some consumers as escargot.[9] Terms such as "garden snail" or "common brown garden snail" are rather meaningless, since they refer to so many types of snails, but they sometimes mean H. aspersa.

  • Cornu aspersum, formerly officially called Helix aspersa Müller, is also known as the French petit gris, "small grey snail", the escargot chagrine, or la zigrinata. The shell of a mature adult has four or five whorls and measures 30 to 45 millimetres (1+18 to 1+34 in) across. It is native to the shores of the Mediterranean and along the coasts of Spain and France. It is found on many British Isles, where the Romans introduced it in the first century AD (some references say it dates to the early Bronze Age). In the early 19th century, the French brought it into California, where it has become a serious pest. These snails are now common throughout the U.S. It was introduced into several Eastern and Gulf states even before 1850, and later introduced into other countries such as Australia, South Africa, New Zealand, Mexico, and Argentina. C. aspersum has a lifespan of 2 to 5 years. This species is more adaptable to different climates and conditions than many snails, and is found in woods, fields, sand dunes, and gardens. This adaptability not only increases C. aspersum's range, but it also makes farming it easier and less risky.
  • Helix pomatia measures about 45 millimetres (1+34 in) across the shell.[10] It also is called the "Roman snail", "apple snail", "lunar", la vignaiola, Weinbergschnecke, escargot de Bourgogne or "Burgundy snail", or "gros blanc. Native over a large part of Europe, it lives in wooded mountains and valleys up to 2,000 metres (6,600 feet) altitude and in vineyards and gardens.[11] The Romans may have introduced it into Britain during the Roman period (AD 43–410).[12] Immigrants introduced it into the U.S. in Michigan and Wisconsin. Many prefer H. pomatia to H. aspersa for its flavor and larger size, as the "escargot par excellence. To date, H. pomatia, however, has not been economically viable for farming.
  • Otala lactea is sometimes called the "vineyard snail", "milk snail", or "Spanish snail".[13] The shell is white with reddish-brown spiral bands, and measures about 26 to 35 millimetres (1 to 1+38 in) in diameter.
  • Iberus alonensis, the Spanish vaqueta or serrana, measures about 30 millimetres (1+18 in) across the shell.
  • Cepaea nemoralis, the "grove snail" or Spanish vaqueta, measures about 25 millimetres (1 in) across the shell.[14] It inhabits Central Europe and was introduced into, and is now naturalized in, many U.S. states, from Massachusetts to California, and from Tennessee to Canada. Its habitat ranges widely from woods to dunes. It mainly eats dead plant material, but it likes nettles and buttercups, and eats dead worms and snails.
  • Cepaea hortensis, the "white-lipped snail", measures about 20 millimetres (34 in) across the shell, which often has distinct dark stripes. It is native to central and northern Europe. This species was introduced into Maine, Massachusetts, and New Hampshire in colonial times, but it never became established in these states. Its habitat varies, but C. hortensis is found in colder and wetter places than C. nemoralis. Their smaller size and some people's opinion that they do not taste as good make C. hortensis and C. nemoralis less popular than the larger European land snails.
  • Otala punctata, called vaqueta in some parts of Spain, measures about 35 millimetres (1+38 in) across the shell.
  • Eobania vermiculata, the vinyala, "mongeta, or xona, measures about 25 millimetres (1 in). It is found in Mediterranean countries and was introduced into Louisiana and Texas.
  • Helix lucorum, commonly called the Turkish snail because of it prevalence in Turkey, measures about 45 millimetres (1+34 in) across the shell. It is found in central Italy and from Yugoslavia through the Crimea to Turkey and around the Black Sea.
  • Helix adanensis comes from around Turkey.
  • Helix aperta measures about 25 millimetres (1 in). Its meat is highly prized. It is native to France, Italy, and other Mediterranean countries, and has become established in California and Louisiana. Sometimes known as the "burrowing snail", it is found above ground only during rainy weather. In hot, dry weather, it burrows 3–6 inches (7.6–15.2 cm) into the ground and becomes dormant until rain softens the soil.
  • Theba pisana, also called the "banded snail" or the "cargol avellanenc", measures about 20 millimetres (34 in) and lives on dry, exposed sites, usually near the sea. Native to Sicily, it has been spread to several European countries, including England. This snail is a serious garden pest and is the "white snail" that California once eradicated using flamethrowers to burn off whole areas. In large numbers, up to 3,000 snails per tree, it can ravage a garden in 24 hours and a citrus or other crop in a few nights.
  • Sphincterochila candidissima or Leucochroa candidissima, the "cargol mongeta or cargol jueu, measures about 20 millimetres (34 in).
  • Lissachatina fulica (formerly Achatina fulica) and other species in the family Achatinidae, giant African snails, can grow up to 326 millimetres (1 ft 34 in) in length. Their native range is south of the Sahara in East Africa. This snail was purposely introduced into India in 1847. An unsuccessful attempt was made to establish it in Japan in 1925. It has been purposely and accidentally transported to other Pacific locations and was inadvertently released in California after World War II, in Hawaii, and later in North Miami, Florida, in the 1970s. In many places, it is a serious agricultural pest that causes considerable crop damage. Also, due to its large size, its slime and fecal material create a nuisance as does the odor that occurs when something like poison bait causes large numbers to die. The U.S. has made considerable effort to eradicate these snails. The U.S. Department of Agriculture has banned the importation and possession of live giant African snails.[15] However, they are still sought after as pets due to the vibrant "tiger stripes" on their shells. Giant African snails can be farmed, but their requirements and their farming methods differ significantly from those of the farming of Helix species.

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Cornu aspersum

Cornu aspersum

Cornu aspersum, known by the common name garden snail, is a species of land snail in the family Helicidae, which includes some of the most familiar land snails. Of all terrestrial molluscs, this species may well be the most widely known. It was classified under the name Helix aspersa for over two centuries, but the prevailing classification now places it in the genus Cornu.

Helix pomatia

Helix pomatia

Helix pomatia, common names the Roman snail, Burgundy snail, or escargot, is a species of large, edible, air-breathing land snail, a pulmonate gastropod terrestrial mollusc in the family Helicidae. It is one of Europe's biggest species of land snail.

Lissachatina fulica

Lissachatina fulica

Lissachatina fulica is a species of large land snail that belongs in the subfamily Achatininae of the family Achatinidae. It is also known as the Giant African land snail. It shares the common name "giant African snail" with other species of snails such as Achatina achatina and Archachatina marginata. This snail species has been considered a significant cause of pest issues around the world. Internationally, it is the most frequently occurring invasive species of snail.

Canning

Canning

Canning is a method of food preservation in which food is processed and sealed in an airtight container. Canning provides a shelf life that typically ranges from one to five years, although under specific circumstances, it can be much longer. A freeze-dried canned product, such as canned dried lentils, could last as long as 30 years in an edible state.

Roman Empire

Roman Empire

The Roman Empire was the post-Republican period of ancient Rome. As a polity, it included large territorial holdings around the Mediterranean Sea in Europe, North Africa, and Western Asia, and was ruled by emperors. From the accession of Caesar Augustus as the first Roman emperor to the military anarchy of the 3rd century, it was a Principate with Italia as the metropole of its provinces and the city of Rome as its sole capital. The Empire was later ruled by multiple emperors who shared control over the Western Roman Empire and the Eastern Roman Empire. The city of Rome remained the nominal capital of both parts until AD 476 when the imperial insignia were sent to Constantinople following the capture of the Western capital of Ravenna by the Germanic barbarians. The adoption of Christianity as the state church of the Roman Empire in AD 380 and the fall of the Western Roman Empire to Germanic kings conventionally marks the end of classical antiquity and the beginning of the Middle Ages. Because of these events, along with the gradual Hellenization of the Eastern Roman Empire, historians distinguish the medieval Roman Empire that remained in the Eastern provinces as the Byzantine Empire.

Otala lactea

Otala lactea

Otala lactea, known as the milk snail or Spanish snail, is a large, edible species of air-breathing land snail, a terrestrial pulmonate gastropod mollusk, in the family Helicidae, the typical snails.

Iberus alonensis

Iberus alonensis

Iberus alonensis is a species of gastropods belonging to the family Helicidae.

Otala punctata

Otala punctata

Otala punctata is a species of air-breathing land snail, a terrestrial pulmonate gastropod mollusk in the family Helicidae.

Eobania vermiculata

Eobania vermiculata

Eobania vermiculata also known as Helix vermiculata, common name the "chocolate-band snail" is a species of large, air-breathing, land snail, a terrestrial pulmonate gastropod mollusk in the family Helicidae, the true snails or typical snails.

Helix lucorum

Helix lucorum

Helix lucorum is a species of large, edible, air-breathing land snail, a terrestrial pulmonate gastropod mollusk in the family Helicidae, the typical snails.

Theba pisana

Theba pisana

Theba pisana, common names the white garden snail, sand hill snail, white Italian snail, Mediterranean coastal snail, and simply just the Mediterranean snail, is an edible species of medium-sized, air-breathing land snail, a terrestrial pulmonate gastropod mollusk in the family Helicidae, the typical snails.

Sphincterochila candidissima

Sphincterochila candidissima

Sphincterochila candidissima is a species of air-breathing land snail, a terrestrial pulmonate gastropod mollusk in the family Sphincterochilidae.

Biology

Understanding of the snail's biology is fundamental to use the right farming techniques. The snail's biology is therefore described here with that in mind.

Anatomy

The anatomy of the edible land snail is described in Land snail.

Lifecycle

General

Snails are hermaphrodites. Although they have both male and female reproductive organs, they must mate with another snail of the same species before they lay eggs. Some snails may act as males one season and as females the next. Other snails play both roles at once and fertilize each other simultaneously. When the snail is large enough and mature enough, which may take several years, mating occurs in the late spring or early summer after several hours of courtship. Sometimes, a second mating occurs in summer. (In tropical climates, mating may occur several times a year. In some climates, snails mate around October and may mate a second time 2 weeks later.) After mating, the snail can store sperm received for up to a year, but it usually lays eggs within a few weeks. Snails are sometimes uninterested in mating with another snail of the same species that originated from a considerable distance away. For example, an H. aspersa from southern France may reject an H. aspersa from northern France.

Helix pomatia

Snails need soil at least 2 inches deep in which to lay their eggs. For H. pomatia, the soil should be at least 3 inches deep to keep out pests such as ants, earwigs, millipedes, etc. Dry soil is not suitable for the preparation of a nest, nor is soil that is too heavy. In clay soil that becomes hard, reproduction rates may decrease because the snails are unable to bury their eggs and the hatchlings have difficulty emerging from the nest. The hatchability of eggs depends on soil temperature, soil humidity, soil composition, etc. Soil consisting of 20% to 40% organic material is good. The soil should be kept at 41 to 50 °F (5 to 10 °C), and is best around 70 °F (21 °C). Soil moisture should be maintained at 80%. One researcher removes eggs immediately after they are deposited, counts them, then keeps them on moist cotton until the eggs hatch and the young start to eat. Snails lose substantial weight by laying eggs. Some do not recover. About one-third of the snails will die after the breeding season.

Helix pomatia eggs measure about 3 mm in diameter and have a calcareous shell and a high yolk content. Eggs are laid in July or August, 2 to 8 weeks after mating, in holes dug out in the ground. (Data varies widely on how long after mating snails lay eggs.) The snail puts its head into the hole or may crawl in until only the top of the shell is visible; then it deposits eggs from the genital opening just behind the head. It takes the snail 1 to 2 days to lay 30 to 50 eggs. Occasionally, the snail will lay about a dozen more a few weeks later. The snail covers the hole with a mixture of the slime it excretes and dirt. This slime, which the snail excretes to help it crawl and to help preserve the moisture in its soft body, is glycoprotein similar to egg white.

Fully developed baby H. pomatia snails hatch about 3 to 4 weeks after the eggs are laid, depending on temperature and humidity. Birds, insects, mice, toads and other predators take a heavy toll on the young snails. The snails eat and grow until the weather turns cold. They then dig a deep hole, sometimes as deep as 1-foot (30 cm), and seal themselves inside their shell and hibernate for the winter. This is a response to both decreasing temperature and shorter hours of daylight. When the ground warms up in spring, the snail emerges and goes on a binge of replacing lost moisture and eating.

Helix aspersa

Helix aspersa eggs are white, spherical, about 3 mm (18 in) in diameter and are laid 5 days to 3 weeks after mating. (Data varies widely due to differences in climate and regional variations in the snails' habitats.) H. aspersa lays an average of 85 eggs in a nest that is 1 to 1+12 inches (2.5 to 3.8 cm) deep. Data varies from 30 to over 120 eggs, but high figures may be from when more than one snail lays eggs in the same nest.

In warm, damp climates, H. aspersa may lay eggs as often as five times from February through October, depending on the weather and region. Mating and egg-laying begin when there are at least 10 hours of daylight and continue until days begin to get shorter. In the United States, longer hours of sunlight that occur when temperatures are still too cold will affect this schedule, but increasing hours of daylight still stimulate egg laying. If warm enough, the eggs hatch in about 2 weeks, or in 4 weeks if cooler. It takes the baby snails several more days to break out of the sealed nest and climb to the surface. In a climate similar to southern California's, H. aspersa matures in about 2 years. In central Italy, H. aspersa hatches and emerges from the soil almost exclusively in the autumn. If well fed and not overcrowded, those snails that hatch at the start of the season will reach adult size and form a lip at the edge of their shell by the following June. If the environment is manipulated to get more early hatchlings, the size and number of snails that mature the following year will increase. In South Africa, some H. aspersa mature in 10 months, and under ideal conditions in a laboratory, some have matured in 6 to 8 months. Most of H. aspersa's reproductive activity takes place in the second year of its life.

Achatina fulica

One giant African snail, Achatina fulica, lays 100 to 400 elliptical eggs that each measure about 5 mm long. Each snail may lay several batches of eggs each year, usually in the wet season. They may lay eggs in holes in the ground like H. pomatia, or lay eggs on the surface of a rocky soil, in organic matter, or at the base of plants. In 10 to 30 days, the eggs hatch releasing snails about 4 mm long. These snails grow up to 10 mm per month. After 6 months, the A. fulica is about 35 mm long and may already be sexually mature. Sexual maturity takes 6 to 16 months, depending on weather and the availability of calcium. This snail lives 5 or 6 years, sometimes as many as 9 years.

Growth

Within the same snail population and under the same conditions, some snails grow faster than others. Some take twice as long to mature. This may help the species survive bad weather, etc., in the wild. However, a snail farmer should obviously select and keep the largest and fastest-maturing snails for breeding stock and sell the smaller snails. By selecting only the largest, the average size of the snail may increase significantly in only a few generations. Most of the differences in growth are probably due to environmental factors, including stocking density. To whatever extent these differences are genetic, though, farmers generally breed large, fast-growing snails instead of small, slower-growing ones.

Several factors can greatly influence the growth of snails, including population density, stress (snails are sensitive to noise, light, vibration, unsanitary conditions, irregular feedings, being touched, etc.), feed, temperature and moisture, and the breeding technology used.

Helix aspersa requires at least 3 to 4% calcium in the soil (or another source of calcium) for good growth. Most snails need more calcium in the soil than H. aspersa. Low calcium intake slows the growth rate and causes the shells to be thinner. Calcium may be set out in a feeding dish or trough so the snails can eat it at will. Food is only one calcium source. Snails may eat paint or attack walls of buildings seeking calcium, and they also eat dirtsoil.

A newly hatched snail's shell size depends on the egg size since the shell develops from the egg's surface membrane. As the snail grows, the shell is added in increments. Eventually, the shell develops a flare or reinforcing lip at its opening. This shows that the snail is now mature; no further shell growth can occur. Growth is measured by shell size, since a snail's body weight fluctuates, even in 100% humidity. The growth rate varies considerably between individuals in each population group. Adult size, which is related to the growth rate, also varies, thus the fastest growers are usually the largest snails. Eggs from larger, healthier snails also tend to grow faster and thus larger.

Dryness inhibits growth and even stops activity. When weather becomes too hot and dry in summer, the snail becomes inactive, seals its shell, and estivates (becomes dormant) until cooler, moister weather returns. Some snails estivate in groups on tree trunks, posts, or walls. They seal themselves to the surface, thus sealing up the shell opening.

Peak snail activity (including feeding and thus growth) occurs a few hours after sunset, when the temperature is lower and the water content (in the form of dew) is higher. During daytime, snails usually seek shelter.

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Land snail

Land snail

A land snail is any of the numerous species of snail that live on land, as opposed to the sea snails and freshwater snails. Land snail is the common name for terrestrial gastropod mollusks that have shells. However, it is not always easy to say which species are terrestrial, because some are more or less amphibious between land and fresh water, and others are relatively amphibious between land and salt water.

Hermaphrodite

Hermaphrodite

In reproductive biology, a hermaphrodite is an organism that has both kinds of reproductive organs and can produce both gametes associated with male and female sexes.

Courtship

Courtship

Courtship is the period wherein some couples get to know each other prior to a possible marriage or committed romantic, de facto relationship. Courtship traditionally may begin after a betrothal and may conclude with the celebration of marriage. A courtship may be an informal and private matter between two people or may be a public affair, or a formal arrangement with family approval. Traditionally, in the case of a formal engagement, it is the role of a male to actively "court" or "woo" a female, thus encouraging her to understand him and her receptiveness to a marriage proposal.

Earwig

Earwig

Earwigs make up the insect order Dermaptera. With about 2,000 species in 12 families, they are one of the smaller insect orders. Earwigs have characteristic cerci, a pair of forcep-like pincers on their abdomen, and membranous wings folded underneath short, rarely used forewings, hence the scientific order name, "skin wings". Some groups are tiny parasites on mammals and lack the typical pincers. Earwigs are found on all continents except Antarctica.

Millipede

Millipede

Millipedes are a group of arthropods that are characterised by having two pairs of jointed legs on most body segments; they are known scientifically as the class Diplopoda, the name derived from this feature. Each double-legged segment is a result of two single segments fused together. Most millipedes have very elongated cylindrical or flattened bodies with more than 20 segments, while pill millipedes are shorter and can roll into a tight ball. Although the name "millipede" derives from the Latin for "thousand feet", no species was known to have 1,000 or more until the discovery of Eumillipes persephone, which can have over 1,300 legs. There are approximately 12,000 named species classified into 16 orders and around 140 families, making Diplopoda the largest class of myriapods, an arthropod group which also includes centipedes and other multi-legged creatures.

Soil moisture

Soil moisture

Soil moisture is the water content of the soil. It can be expressed in terms of volume or weight. Soil moisture measurement can be based on in situ probes or remote sensing methods.

Calcareous

Calcareous

Calcareous is an adjective meaning "mostly or partly composed of calcium carbonate", in other words, containing lime or being chalky. The term is used in a wide variety of scientific disciplines.

Yolk

Yolk

Among animals which produce eggs, the yolk is the nutrient-bearing portion of the egg whose primary function is to supply food for the development of the embryo. Some types of egg contain no yolk, for example because they are laid in situations where the food supply is sufficient or because the embryo develops in the parent's body, which supplies the food, usually through a placenta. Reproductive systems in which the mother's body supplies the embryo directly are said to be matrotrophic; those in which the embryo is supplied by yolk are said to be lecithotrophic. In many species, such as all birds, and most reptiles and insects, the yolk takes the form of a special storage organ constructed in the reproductive tract of the mother. In many other animals, especially very small species such as some fish and invertebrates, the yolk material is not in a special organ, but inside the egg cell.

Glycoprotein

Glycoprotein

Glycoproteins are proteins which contain oligosaccharide chains covalently attached to amino acid side-chains. The carbohydrate is attached to the protein in a cotranslational or posttranslational modification. This process is known as glycosylation. Secreted extracellular proteins are often glycosylated.

Sexual maturity

Sexual maturity

Sexual maturity is the capability of an organism to reproduce. In humans, it is related to both puberty and adulthood. However, puberty is the process of biological sexual maturation, while the concept of adulthood is generally based on broader cultural definitions.

Calcium

Calcium

Calcium is a chemical element with the symbol Ca and atomic number 20. As an alkaline earth metal, calcium is a reactive metal that forms a dark oxide-nitride layer when exposed to air. Its physical and chemical properties are most similar to its heavier homologues strontium and barium. It is the fifth most abundant element in Earth's crust, and the third most abundant metal, after iron and aluminium. The most common calcium compound on Earth is calcium carbonate, found in limestone and the fossilised remnants of early sea life; gypsum, anhydrite, fluorite, and apatite are also sources of calcium. The name derives from Latin calx "lime", which was obtained from heating limestone.

Dew

Dew

Dew is water in the form of droplets that appears on thin, exposed objects in the morning or evening due to condensation.

Values of the snail

Nutritional value

The nutrient composition of raw snails (per 100 g of edible portion), according to information from the nutrient databank of France, is:

Biochemical content

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Food energy

Food energy

Food energy is chemical energy that animals derive from their food to sustain their metabolism, including their muscular activity.

Calorie

Calorie

The calorie is a unit of energy that originated from the obsolete caloric theory of heat. For historical reasons, two main definitions of "calorie" are in wide use. The large calorie, food calorie, dietary calorie, or kilogram calorie was originally defined as the amount of heat needed to raise the temperature of one kilogram of water by one degree Celsius. The small calorie or gram calorie was defined as the amount of heat needed to cause the same increase in one gram of water. Thus, 1 large calorie is equal to 1000 small calories.

Protein

Protein

Proteins are large biomolecules and macromolecules that comprise one or more long chains of amino acid residues. Proteins perform a vast array of functions within organisms, including catalysing metabolic reactions, DNA replication, responding to stimuli, providing structure to cells and organisms, and transporting molecules from one location to another. Proteins differ from one another primarily in their sequence of amino acids, which is dictated by the nucleotide sequence of their genes, and which usually results in protein folding into a specific 3D structure that determines its activity.

Dietary fiber

Dietary fiber

Dietary fiber or roughage is the portion of plant-derived food that cannot be completely broken down by human digestive enzymes. Dietary fibers are diverse in chemical composition, and can be grouped generally by their solubility, viscosity, and fermentability, which affect how fibers are processed in the body. Dietary fiber has two main components: soluble fiber and insoluble fiber, which are components of plant-based foods, such as legumes, whole grains and cereals, vegetables, fruits, and nuts or seeds. A diet high in regular fiber consumption is generally associated with supporting health and lowering the risk of several diseases. Dietary fiber consists of non-starch polysaccharides and other plant components such as cellulose, resistant starch, resistant dextrins, inulin, lignins, chitins, pectins, beta-glucans, and oligosaccharides.

Fat

Fat

In nutrition, biology, and chemistry, fat usually means any ester of fatty acids, or a mixture of such compounds, most commonly those that occur in living beings or in food.

Magnesium

Magnesium

Magnesium is a chemical element with the symbol Mg and atomic number 12. It is a shiny gray metal having a low density, low melting point and high chemical reactivity. Like the other alkaline earth metals it occurs naturally only in combination with other elements and it almost always has an oxidation state of +2. It reacts readily with air to form a thin passivation coating of magnesium oxide that inhibits further corrosion of the metal. The free metal burns with a brilliant-white light. The metal is obtained mainly by electrolysis of magnesium salts obtained from brine. It is less dense than aluminium and is used primarily as a component in strong and lightweight alloys that contain aluminium.

Calcium

Calcium

Calcium is a chemical element with the symbol Ca and atomic number 20. As an alkaline earth metal, calcium is a reactive metal that forms a dark oxide-nitride layer when exposed to air. Its physical and chemical properties are most similar to its heavier homologues strontium and barium. It is the fifth most abundant element in Earth's crust, and the third most abundant metal, after iron and aluminium. The most common calcium compound on Earth is calcium carbonate, found in limestone and the fossilised remnants of early sea life; gypsum, anhydrite, fluorite, and apatite are also sources of calcium. The name derives from Latin calx "lime", which was obtained from heating limestone.

Iron

Iron

Iron is a chemical element with symbol Fe and atomic number 26. It is a metal that belongs to the first transition series and group 8 of the periodic table. It is, by mass, the most common element on Earth, just ahead of oxygen, forming much of Earth's outer and inner core. It is the fourth most common element in the Earth's crust, being mainly deposited by meteorites in its metallic state, with its ores also being found there.

Vitamin C

Vitamin C

Vitamin C is a water-soluble vitamin found in citrus and other fruits and vegetables, also sold as a dietary supplement and as a topical "serum" ingredient to treat melasma and wrinkles on the face. It is used to prevent and treat scurvy. Vitamin C is an essential nutrient involved in the repair of tissue, the formation of collagen, and the enzymatic production of certain neurotransmitters. It is required for the functioning of several enzymes and is important for immune system function. It also functions as an antioxidant. Most animals are able to synthesize their own vitamin C. However, apes and monkeys, most bats, some rodents, and certain other animals must acquire it from dietary sources.

Snail farming

Successful snail culture requires the correct equipment and supplies, including snail pens or enclosures; devices for measuring humidity (hygrometer), temperature (thermometer), soil moisture (soil moisture sensor), and light (in foot candles); a weight scale and an instrument to measure snail size; a kit for testing soil contents; and a magnifying glass to see the eggs. Equipment to control the climate (temperature and humidity), to regulate water (e.g., a sprinkler system to keep the snails moist and a drainage system), to provide light and shade, and to kill or keep out pests and predators may also be needed. Some horticultural systems such as artificial lighting systems and water sprinklers may be adapted for snail culture. Better results are obtained if snails of the same kind and generation are used. Some recommend putting the hatchlings in another pen.

Four systems of snail farms can be distinguished:

  • Outdoor pens
  • In buildings with a controlled climate
  • In closed systems such as plastic tunnel houses or "greenhouses"
  • In addition, snails may breed and hatch inside in a controlled environment and then (after 6 to 8 weeks) may be placed in outside pens to mature.

Key factors to successful snail farming

Hygiene

Good hygiene can prevent the spread of disease and otherwise improve the health and growth rate of snails. Food is replaced daily to prevent spoilage. Earthworms added to the soil helps keep the pen clean.

Parasites, nematodes, trematodes, fungi, and microarthropods can attack snails, and such problems can spread rapidly when snail populations are dense. The bacterium Pseudomonas aeruginosa causes intestinal infections that can spread rapidly in a crowded snail pen.

Possible predators include rats, mice, moles, skunks, weasels, birds, frogs and toads, lizards, walking insects (e.g., some beetle and cricket species), some types of flies, centipedes, and even certain carnivorous snail species, such as Strangesta capillacea.

Population density

Population density also affects successful snail production. Pens should contain no more than six to eight fair-sized snails per square foot (65 to 86/m2), or about four large H. pomatias per square foot (43/m2); or figure one kilogram per square metre (0.20 lb/sq ft), which automatically compensates for the size of the snails. To encourage breeding, best results will occur with not more than eight snails per square metre (0.74/sq ft). Some sources say that, for H. pomatia to breed, 2 to 4 snails per square metre (0.19 to 0.37/sq ft) is the maximum.

Snails tend not to breed when packed too densely or when the slime in the pen accumulates too much. The slime apparently works like a pheromone and suppresses reproduction. On the other hand, snails in groups of about 100 seem to breed better than when only a few snails are confined together. Perhaps they have more potential mates from which to choose. Snails in a densely populated area grow more slowly even when food is abundant, and they also have a higher mortality rate. These snails then become smaller adults who lay fewer clutches of eggs, have fewer eggs per clutch, and the eggs have a lower hatch rate. Smaller adult snails sell for less. Dwarfing is quite common in snail farming and is attributable mainly to rearing conditions rather than heredity factors. Crowding snails is false economy. A recommended rate for H. aspersa is not more than 13 pound per square foot (1.6 kg/m2) of soil surface for snails that weigh more than 1 gram (0.035 oz) for snails that weigh less.

Feeding

The feeding season is April through October, (or may vary with the local climate), with a "rest period" during the summer. Do not place food in one small clump so that there is not enough room for all the snails to get to it. Snails eat solid food by rasping it away with their radula. Feeding activity depends on the weather, and snails may not necessarily feed every day. Evening irrigation in dry weather may encourage feeding since the moisture makes it easier for the snails to move about.

Put the breeding snails in the breeding pens in April or early May. Feed until mid-June when mating begins and the snails stop feeding. Snails resume eating after they lay eggs. Once snails have laid their eggs, the adult snails can be removed. This leaves more food and less crowding for the hatchlings.

Snails of the same species collected from different regions may have different food preferences. Some foods that snails eat are: Alyssum, fruit and leaves of apple, apricot, artichoke (a favorite), aster, barley, beans, bindweed, California boxwood, almost any cabbage variety, camomile, carnation, carrot, cauliflower, celeriac (root celery), celery, ripe cherries, chive, citrus, clover, cucumbers (a favorite snail food), dandelion, elder, henbane, hibiscus, hollyhock, kale, larkspur (Consolida or Delphinium genera), leek, lettuce (liked, and makes good snails), lily, magnolia, mountain ash, mulberry, chrysanthemum, nasturtium, nettle, nightshade berries, oats, onion greens, pansy, parsley, peach, ripe pears, peas, petunia, phlox, plum, potatoes (raw or cooked), pumpkins, radish, rape, rose, sorrel, spinach, sweet pea, thistle, tomatoes (well liked), turnip, wheat, yarrow, zinnia. They will eat sweet lupines, but will reject bitter lupines and other plants with high quinolizidine alkaloids. Snails also avoid plants that produce other defensive chemicals, defensive stem hairs, etc.

Snails usually prefer juicy leaves and vegetables over dry ones. If snails are fed vegetable trimmings, damaged fruit, and cooked potatoes, uneaten food must be promptly removed as it will quickly spoil. Bran may be supplied that is wet or sprinkled dry over leafy vegetables. The diet may consist of 20% wheat bran while 80% is fruit and vegetable material. Some growers use oats, corn meal, soybean meal, or chicken mash. Laying mash provides calcium, as does crushed oyster shells. Snails also may eat materials such as cardboard (but do not purposely feed it to them); they can eat through shipping cartons and escape. Snails may sometimes eat, within a 24-hour period, food equal to 10%, and occasionally as much as 20%, of their body weight. Active snails deprived of food will lose more than one-third of their weight before they starve to death—a process that takes 8 to 12 weeks. Estivating snails can survive much longer.

Supply calcium at least once a week if it is not available in the soil. It should not contain harmful salts or be so alkaline as to burn the snails. Mix calcium with wet bran or mashed potatoes and serve on a pan; this will keep any leftover food from rotting on the ground.

Some researchers use chicken mash for feed. A plastic pipe can be cut in half lengthwise to make two troughs which can be used as feeders for mash. Mix laying mash (used for egg-producing hens) into the feed to provide calcium for the snails' shells. Commercial chicken feeding mash is around 16% to 17% protein, from fish meal and meat meal, making it good for growing snails. Supplying mash to hatchlings might reduce cannibalism. Two feeds that snails like and that promote good growth are: (A) broiler finisher mash consisting of 7% broiler concentrate, 58% corn, 16% soya, 18% sorghum, 7% limestone flour (40% Ca); and (B) chicken feed (pellets) for layers consisting of 5% layer concentrate, 10%, corn, 15% soya, 20% sorghum, 44% barley, 6% limestone flour (40% Ca).

Pellets are fine for larger snails, but mash is better for younger ones. Pellets should be partially crushed if fed to young snails. Snails do not grow well if rabbit pellets are their primary diet. Snails show a distinct preference for moist feed. Easy access to enough water must be ensured if snails are fed dry mash.

Be sure to frequently clean the feed and water dishes. The amount of feed a snail eats depends very much on air humidity and on the availability of drinking water. Clean drinking water can be served in a shallow container to reduce the risk of the snail drowning. Some types of chicken waterers may be suitable. Other factors (e.g., temperature, light intensity, food preferences versus food supplied, etc.) also affect feeding. A compromise, until the optimum feed is found, is to feed half green vegetable material and half chicken feed/grain/animal protein.

Young of H. aspersa readily eats milk powder. Its rapid rate of assimilation promotes rapid growth.

Climate

A mild climate 15–25 °C (59–77 °F) with high humidity (75% to 95%) is best for snail farming, though most varieties can stand a wider range of temperatures. The optimal temperature is 21 °C (70 °F) for many varieties. When the temperature falls below 7 °C (45 °F), snails hibernate. Under 12 °C (54 °F) the snails are inactive, and under 10 °C (50 °F), all growth stops. When the temperature rises much above 27 °C (81 °F) or conditions become too dry, snails estivate. Wind is bad for snails because it speeds up moisture loss, and snails must retain moisture.

Snails thrive in damp but not waterlogged environments and thus a well-draining soil is required. Research indicates that water content around 80% of the carrying capacity of the soil and air humidity over 80% (during darkness) are the most favorable conditions. Many farmers use mist-producing devices to maintain proper moisture in the air and/or soil.[16] Also, if the system contains alive vegetation, the leaves are to be periodically wet.

Soil

Snails dig in soil and ingest it. Good soil favors snail growth and provides some of their nutrition. Lack of access to good soil may cause fragile shells even when the snails have well-balanced feed; the snails' growth may lag far behind the growth of other snails on good soil. Snails often eat feed, then go eat soil. Sometimes, they eat only one or the other.

A well-functional soil should have these characteristics:

  • Containing neither too much sand nor too much clay, as snails strive to dig into hard clay and sand dries out easily
  • A 20–40% organic matter content - organic matter enhances cation exchange capacity of calcium and magnesium, which in turn stimulates growth.
  • pH around 7
  • Adequate calcium, the primary constituent of shells (up to 98%) - a common way to introduce calcium is to add ground limestone at a suggested concentration of 0.5 kg/m2 (0.1 lb/sq ft). Calcium may also be set out in a feeding dish or trough so the snails can eat it at will. More advanced techniques involve the addition of polyacrylamide with the following concentration: 12.5 cm3 of a 160 g M.A./one preparation in 0.25 L/kg of water in dry soil. Such stabilization treatment helps the soil structure resist washing and allows regular cleaning without destroying the crumb structure of the soil that is beneficial for egg laying.

Soil care: A farmer must find a way to prevent the soil from becoming fouled with mucus and droppings and also tackle undesirable chemical changes that may occur in time.

Soil mix suggestions:

Phases in snail farming

Some who raise H. aspersa separate the five stages: reproduction, hatching, young, fattening, and final fattening.

Depending on the scale and sophistication of a snail farm, it will contain some or all of below described sections which may or may not be merged with one and another. Each section has its particular values for the key factors to successful snail farming, described above.

Hibernation

For future reproducers it is mandatory to hibernate 3 months.

Breeding

Most breeders allow the snails to mate with one another on their own. If snails are kept in ideal conditions, breeding will occur at higher rates and have more success.

Hatchery and nursery

When the snails have laid their eggs, the pots are put in a nursery where the eggs will hatch. The young snails are kept in the nursery for about 6 weeks, and then moved to a separate pen, as young snails do best if kept with other snails of similar size. Eight hours of daylight is optimal for young snails.

Baby snails are fed on tender lettuce leaves (Boston type, but head type is probably also good).

Cannibalism by hatchlings

The first snails to hatch eat the shells of their eggs. This gives them calcium needed for their shells. They may then begin eating unhatched eggs. If the snail eggs are kept at the optimum temperature, 68 °F (20 °C) (for some varieties), and if none of the eggs lose moisture, most eggs will hatch within 1 to 3 days of each other. Cannibalism also will be low. If hatching extends over a longer period, cannibalism may increase. Some eggs eaten are eggs that were not fertile or did not develop properly, but sometimes, properly developing embryos might be eaten. A high density of "clutches" of egg masses increases the rate of cannibalism, as other nearby egg masses are more likely to be found and eaten. Snail egg has 12 to 20 times the protein of lettuce. The protein helps the baby snails start developing quickly and be healthier. Snail egg is an excellent starter food for newly hatched snails, but they tend to only eat eggs of their own species.

Fattening/growing

In this section, the snails are grown from juvenile to mature size.

Fattening pens can be outside or in a greenhouse. High summer temperatures and insufficient moisture cause dwarfing and malformations of some snails. This is more of a problem inside greenhouses if the sun overheats the building. A sprinkler system (e.g., a horticultural system or common lawn sprinklers) can supply moisture. Make sure excess water can drain.

Fattening pens may contain 2-by-3-foot (0.61 by 0.91 m) pieces (or other convenient size) of heavy plastic sheets, hung from boards resting on a rack that lets the tips of the plastic sheets just touch the ground. The plastic sheets are about 4 inches (10 cm) apart. The sheets give the snails a resting and hiding place. Feeders may be located on the rack that supports the plastic sheets.

A layer of coarse sand and topsoil with earthworms is placed on the fattening pen's bottom. The worms help clean up the snail droppings.

Snails that hatched the previous summer can be put in a chilled room for hibernation over winter. About 1 April, (adjusted for local climate), they are moved to the final fattening pen. In cases where there are several fattening pens, the smaller snails are placed in one, medium in another, large in another. One-third of a pound of H. aspersa snails require one square foot of pen. Snails lose weight when they estivate in summer, therefore some growers do not stock pens by weight but by count. For H. aspersa, 10 to 12 snails per square foot is the maximum.

Harvest and purging

Snails are mature when a lip forms at the opening of their shell. Before they mature, their shells are more easily broken, making them undesirable. For H. aspersa, commercial weight is 8 grams or larger.

The fastest, largest, and healthy snails are selected for next-generation breeders. This is typically around 5% of the harvest. The remainder goes for sales.

Snail eggs may also be harvested and processed to produce snail caviar,[17] but in order to do so systematically, special breeding units are created enhancing easy harvest of the eggs.

Types of farms, or sections thereof

Open air farms

Enclosures for snails are usually long and thin instead of square. This allows the workers to walk around (without harming the snails) and reach into the whole pen. The enclosure may be a trough with sides made of wood, block, fiber cement sheets, or galvanized sheet steel. Cover it with screen or netting. The covering confines the snails and keeps out birds and other predators. Fences or walls are usually 2 feet (60 cm) high plus at least 5 inches (13 cm) into the ground. Fencing made of galvanized metal or hard-plastic sheets helps keep out some predators. A cover will protect against heavy rain. Shade (which may be a fine mesh screen) on warm winter days helps keep the snails dormant. 5 mm mesh or finer is used for pen screens or fences. Pens containing baby snails need a finer mesh.

Snails need hiding places, especially during the warm daytime. Plastic soil drainage pipes from the local garden center can be split in two lengthwise, and stacked one layer one way and the next layer at a right angle, providing shelter and also increasing by 50% the number of snails that can live in the pen.

The bottom of the enclosure, if it is not the ground or trays of dirt, needs be a surface more solid than screening. A snail placed in a wire-mesh-bottom pen will keep crawling, trying to get off the wires and onto solid, more comfortable ground.

Garden farms

An alternate method is to make a square pen with a 10-foot (3.0 m)-square garden in it. Plant about six crops, e.g., nettles and artichokes, inside the pen. The snails will choose what they want to eat. If it has not rained, turn sprinklers on for about 15 minutes at dusk, unless the snails are dormant. A disadvantage to this method is that, if the snails are not mature at the end of the year, it is difficult to replant fresh plant crops in the pens.

Plastic tunnels make cheap, easy snail enclosures, but it is difficult to regulate heat and humidity. The tunnel will be 10 to 20 °F (6 to 11 °C) warmer than the outside, and snails become dormant when the temperature climbs above 80 °F (27 °C).

Indoor farms

When snails are raised indoors under controlled conditions, reproduction varies according to the geographic origin of the breeding stock. For example, one researcher found that H. aspersa snails from Brittany seem to do better indoors than snails from another region. To breed snails indoors, the temperature needs to be kept at 70 °F (21 °C) and the relative humidity at 80% to 90%; some sources say 95%. Another source recommends 75% humidity by day and 95% at night. The Center for Heliciculture once recommended 65–75% humidity during the day and 85–95% at night at 68 °F (20 °C). In any event, avoid humidity higher than 95% (some say 90%) for any length of time. Excessive humidity can kill snails. Optimum temperature and relative humidity depend on several things, including the snail variety and even where breeding stock was gathered. For H. aspersa, the optimum temperature for hatching eggs seems to be 68 °F (20 °C) at 100% relative humidity. The second best temperature/humidity combination depends on where the snails came from and results can drop drastically to 0% hatching at 17 °C (63 °F) and 100% humidity. The soil must not be kept wet when the humidity is maintained at 100%, as the eggs will absorb water, swell up, and burst.

Fluorescent lamps can be used to give artificial daylight. Different snails respond to day length in different ways. The ratio of light to darkness influences activity, feeding, and mating and egg-laying. Eighteen or more hours of light apparently stimulate H. aspersa growth, while less than 12 hours inhibit it. Some snail species may associates the long hours of light with the start of summer—the peak growing season. Eighteen hours of daylight also appear optimal for breeding (mating and egg laying), but snails will breed in darkness.

Snails can be bred in boxes or cages stacked several units high. An automatic sprinkler system can be used to provide moisture. Breeding cages need a feed trough and a water trough. Plastic trays a couple of inches deep are adequate; deeper water troughs increase the chance of snails drowning in them. Trays can be set on a bed of small gravel. Small plastic pots, e.g., flower pots about 3 inches (7.6 cm) deep, can be filled with sterilized dirt (or a loamy pH neutral soil) and set in the gravel to give the snails a place to lay their eggs. After the snails lay eggs each pot is replaced. (Set one pot inside another so that one can be easily lifted without shifting the gravel.)

In a typical example, the breeding box has concrete sides, soil with earthworms (to cleanse the soil) on the bottom, vegetation, curved tiles to provide shelter, feeders, and a chicken waterer. Mosquito netting or screening covers the top. These breeding boxes may be outside, or better results may be obtained when the boxes are inside a greenhouse—as long as the greenhouse does not get too hot or too dry. One researcher reported that in outdoor boxes, each breeder snail had about seven young. In greenhouses, each breeder snail had about 9 to 12 young. The researcher felt that under better weather conditions than those he had that year, each adult breeder snail would have produced 15 young snails.

Systems used in the farm

Fogging system

A sprinkler system ensures moisture when needed and can be turned on at sunset rather than earlier in the day, when the moisture may drive snails out into hot sunshine. Temperature and humidity can be monitored using a thermometer and a hygrometer.

Anti-escape barriers

Except for a box that is closed on all sides, and which is not economic for large scale snail farming, there is no anti-escape barrier that is 100% escape proof. The following methods are just a few systems, each of which has a varying degree of success:

  • in an open pen, the top of the fences curve inward in a half circle; this works to confine the vineyard snail. However, H. aspersa can escape from such an open pen.
  • Electric fence. [The electric fence top has two or more thin wires that are 2 to 4 mm apart. Each wire carries the opposite charge of the wire next to it. A battery or transformer is used to supply 4 to 12 volts to the wire. A snail will get a mild shock and retract when it crawls over a wire and touches a second wire.]
  • Bend the fence top inward into a sharp "V" shape with about a 20-degree angle. The snail's shell will hit the bent-back part of the screen before the snail can reach up and start crawling on it. This blocks the snail, and the angled screen automatically compensates for the size of the snail.
  • Suitable for solid wall enclosures, is to attach to the wall a horizontal piece of screen that projects inward several inches over the enclosure. The screen is made with material such as nylon monofilament that is moderately stiff and springy yet easily flexible. On the inside edge of the screen, the cross fibers are removed, producing a fringe several inches wide. When a snail crawls on the underside of the screen and moves out onto the fringe, his weight pulls several individual fibers down. One by one, another fiber gets away from the snail and springs back up out of reach. Eventually, the snail is dangling by a thread. The snail then falls, because the surface area is not big enough to crawl on.
  • Because snails usually will not cross a copper band, another solution is to top the fence with a copper band at least 3 inches (10 cm) wide. The band can be bent so that part of it faces inward and is parallel to the pen floor. The band must not be placed very close to the ground, because rain may wash soil against the copper and leave a residue that may enable the snail to cross it. The bottom of the fence must be buried deep enough into the ground so that the snails cannot dig under it.

Discover more about Snail farming related topics

Hygrometer

Hygrometer

A hygrometer is an instrument which measures the humidity of air or some other gas: that is, how much water vapor it contains. Humidity measurement instruments usually rely on measurements of some other quantities such as temperature, pressure, mass, a mechanical or electrical change in a substance as moisture is absorbed. By calibration and calculation, these measured quantities can lead to a measurement of humidity. Modern electronic devices use the temperature of condensation, or they sense changes in electrical capacitance or resistance to measure humidity differences. A crude hygrometer was invented by Leonardo da Vinci in 1480. Major leaps came forward during the 1600s; Francesco Folli invented a more practical version of the device, while Robert Hooke improved a number of meteorological devices including the hygrometer. A more modern version was created by Swiss polymath Johann Heinrich Lambert in 1755. Later, in the year 1783, Swiss physicist and Geologist Horace Bénédict de Saussure invented the first hygrometer using human hair to measure humidity.

Humidity

Humidity

Humidity is the concentration of water vapor present in the air. Water vapor, the gaseous state of water, is generally invisible to the human eye. Humidity indicates the likelihood for precipitation, dew, or fog to be present.

Greenhouse

Greenhouse

A greenhouse is a structure with walls and roof made chiefly of transparent material, such as glass, in which plants requiring regulated climatic conditions are grown.These structures range in size from small sheds to industrial-sized buildings. A miniature greenhouse is known as a cold frame. The interior of a greenhouse exposed to sunlight becomes significantly warmer than the external temperature, protecting its contents in cold weather.

Parasitism

Parasitism

Parasitism is a close relationship between species, where one organism, the parasite, lives on or inside another organism, the host, causing it some harm, and is adapted structurally to this way of life. The entomologist E. O. Wilson has characterised parasites as "predators that eat prey in units of less than one". Parasites include single-celled protozoans such as the agents of malaria, sleeping sickness, and amoebic dysentery; animals such as hookworms, lice, mosquitoes, and vampire bats; fungi such as honey fungus and the agents of ringworm; and plants such as mistletoe, dodder, and the broomrapes.

Nematode

Nematode

The nematodes or roundworms constitute the phylum Nematoda, with plant-parasitic nematodes also known as eelworms. They are a diverse animal phylum inhabiting a broad range of environments. Less formally, they are categorized as helminths, but are taxonomically classified along with arthropods, tardigrades and other moulting animals in the clade Ecdysozoa, and unlike flatworms, have tubular digestive systems with openings at both ends. Like tardigrades, they have a reduced number of Hox genes, but their sister phylum Nematomorpha has kept the ancestral protostome Hox genotype, which shows that the reduction has occurred within the nematode phylum.

Arthropod

Arthropod

Arthropods are invertebrate animals with an exoskeleton, a segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and cuticle made of chitin, often mineralised with calcium carbonate. The arthropod body plan consists of segments, each with a pair of appendages. Arthropods are bilaterally symmetrical and their body possesses an external skeleton. In order to keep growing, they must go through stages of moulting, a process by which they shed their exoskeleton to reveal a new one. Some species have wings. They are an extremely diverse group, with up to 10 million species.

Pseudomonas aeruginosa

Pseudomonas aeruginosa

Pseudomonas aeruginosa is a common encapsulated, gram-negative, aerobic–facultatively anaerobic, rod-shaped bacterium that can cause disease in plants and animals, including humans. A species of considerable medical importance, P. aeruginosa is a multidrug resistant pathogen recognized for its ubiquity, its intrinsically advanced antibiotic resistance mechanisms, and its association with serious illnesses – hospital-acquired infections such as ventilator-associated pneumonia and various sepsis syndromes.

Rat

Rat

Rats are various medium-sized, long-tailed rodents. Species of rats are found throughout the order Rodentia, but stereotypical rats are found in the genus Rattus. Other rat genera include Neotoma, Bandicota and Dipodomys.

Mouse

Mouse

A mouse is a small rodent. Characteristically, mice are known to have a pointed snout, small rounded ears, a body-length scaly tail, and a high breeding rate. The best known mouse species is the common house mouse. Mice are also popular as pets. In some places, certain kinds of field mice are locally common. They are known to invade homes for food and shelter.

Mole (animal)

Mole (animal)

Moles are small mammals adapted to a subterranean lifestyle. They have cylindrical bodies, velvety fur, very small, inconspicuous eyes and ears, reduced hindlimbs, and short, powerful forelimbs with large paws adapted for digging.

Bird

Bird

Birds are a group of warm-blooded vertebrates constituting the class Aves, characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweight skeleton. Birds live worldwide and range in size from the 5.5 cm (2.2 in) bee hummingbird to the 2.8 m common ostrich. There are about ten thousand living species, more than half of which are passerine, or "perching" birds. Birds have wings whose development varies according to species; the only known groups without wings are the extinct moa and elephant birds. Wings, which are modified forelimbs, gave birds the ability to fly, although further evolution has led to the loss of flight in some birds, including ratites, penguins, and diverse endemic island species. The digestive and respiratory systems of birds are also uniquely adapted for flight. Some bird species of aquatic environments, particularly seabirds and some waterbirds, have further evolved for swimming.

Frog

Frog

A frog is any member of a diverse and largely carnivorous group of short-bodied, tailless amphibians composing the order Anura. The oldest fossil "proto-frog" Triadobatrachus is known from the Early Triassic of Madagascar, but molecular clock dating suggests their split from other amphibians may extend further back to the Permian, 265 million years ago. Frogs are widely distributed, ranging from the tropics to subarctic regions, but the greatest concentration of species diversity is in tropical rainforest. Frogs account for around 88% of extant amphibian species. They are also one of the five most diverse vertebrate orders. Warty frog species tend to be called toads, but the distinction between frogs and toads is informal, not from taxonomy or evolutionary history.

Processing/transforming snails

Snails can be processed industrially (typically in 'factories') and as a craft (typically in 'kitchens'). Industrial processing of snails risks significant drop of the snail's quality, and with relatively high loss of material. The economies of scale that go with industrial processing, though, allows for profitable compensation. Processing by individual craftsmanship allows for much lower production, but product quality typically remains high.

Market developments

Ukraine

In 2015, the first snail farm opened in Ukraine. Production was, and remains, almost entirely for export, there being no consumer market for snails in the country. Production (in tonnes) was: 93 in 2018; 200–300 in 2019; and 1,000 in 2020, when the country had 400 farms. Exports were decimated in 2020, however, by lockdowns related to the COVID-19 pandemic.[18]

United States

U.S. imports of snails were worth more than $4.5 million in 1995 and came from 24 countries. This includes preserved or prepared snails and snails that are live, fresh, chilled, or frozen. Major exporters to the U.S. are France, Indonesia, Greece and China. The U.S. exported live, fresh, chilled, or frozen snails worth $55,000 to 13 countries; most were shipped to Japan, the Netherlands, and the United Kingdom. Individual statistics are not available for U.S. exports of prepared or processed snails from the U.S. Department of Commerce.

France

The COVID-19 pandemic wiped out almost all sales in France in 2020.[19] This was especially due to the cancelling of New Year's Eve, which comprises 70% of annual sales normally.[19]

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COVID-19 pandemic

COVID-19 pandemic

The COVID-19 pandemic, also known as the coronavirus pandemic, is an ongoing global pandemic of coronavirus disease 2019 (COVID-19) caused by severe acute respiratory syndrome coronavirus 2 (SARS-CoV-2). The novel virus was first identified in an outbreak in the Chinese city of Wuhan in December 2019. Attempts to contain it there failed, allowing the virus to spread to other areas of Asia and later worldwide. The World Health Organization (WHO) declared the outbreak a public health emergency of international concern on 30 January 2020, and a pandemic on 11 March 2020. As of 10 March 2023, the pandemic had caused more than 676 million cases and 6.88 million confirmed deaths, making it one of the deadliest in history.

France

France

France, officially the French Republic, is a country located primarily in Western Europe. It also includes overseas regions and territories in the Americas and the Atlantic, Pacific and Indian Oceans, giving it one of the largest discontiguous exclusive economic zones in the world. Its metropolitan area extends from the Rhine to the Atlantic Ocean and from the Mediterranean Sea to the English Channel and the North Sea; overseas territories include French Guiana in South America, Saint Pierre and Miquelon in the North Atlantic, the French West Indies, and many islands in Oceania and the Indian Ocean. Its eighteen integral regions span a combined area of 643,801 km2 (248,573 sq mi) and had a total population of over 68 million as of January 2023. France is a unitary semi-presidential republic with its capital in Paris, the country's largest city and main cultural and commercial centre; other major urban areas include Marseille, Lyon, Toulouse, Lille, Bordeaux, and Nice.

Indonesia

Indonesia

Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania between the Indian and Pacific oceans. It consists of over 17,000 islands, including Sumatra, Java, Sulawesi, and parts of Borneo and New Guinea. Indonesia is the world's largest archipelagic state and the 14th-largest country by area, at 1,904,569 square kilometres. With over 275 million people, Indonesia is the world's fourth-most populous country and the most populous Muslim-majority country. Java, the world's most populous island, is home to more than half of the country's population.

Greece

Greece

Greece, officially the Hellenic Republic, is a country in Southeast Europe. It is situated on the southern tip of the Balkan Peninsula, and is located at the crossroads of Europe, Asia, and Africa. Greece shares land borders with Albania to the northwest, North Macedonia and Bulgaria to the north, and Turkey to the northeast. The Aegean Sea lies to the east of the mainland, the Ionian Sea to the west, and the Sea of Crete and the Mediterranean Sea to the south. Greece has the longest coastline on the Mediterranean Basin, featuring thousands of islands. The country consists of nine traditional geographic regions, and has a population of approximately 10.4 million. Athens is the nation's capital and largest city, followed by Thessaloniki and Patras.

China

China

China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and borders fourteen countries by land, the most of any country in the world, tied with Russia. With an area of approximately 9.6 million square kilometres (3,700,000 sq mi), it is the world's third largest country by total land area. The country consists of 22 provinces, five autonomous regions, four municipalities, and two special administrative regions. The national capital is Beijing, and the most populous city and largest financial center is Shanghai.

Japan

Japan

Japan is an island country in East Asia. It is situated in the northwest Pacific Ocean and is bordered on the west by the Sea of Japan, extending from the Sea of Okhotsk in the north toward the East China Sea, Philippine Sea, and Taiwan in the south. Japan is a part of the Ring of Fire, and spans an archipelago of 14,125 islands covering 377,975 square kilometers (145,937 sq mi); the five main islands are Hokkaido, Honshu, Shikoku, Kyushu, and Okinawa. Tokyo is the nation's capital and largest city, followed by Yokohama, Osaka, Nagoya, Sapporo, Fukuoka, Kobe, and Kyoto.

Netherlands

Netherlands

The Netherlands, informally Holland, is a country located in northwestern Europe with overseas territories in the Caribbean. It is the largest of four constituent countries of the Kingdom of the Netherlands. The Netherlands consists of twelve provinces; it borders Germany to the east, and Belgium to the south, with a North Sea coastline to the north and west. It shares maritime borders with the United Kingdom, Germany and Belgium in the North Sea. The country's official language is Dutch, with West Frisian as a secondary official language in the province of Friesland. Dutch, English and Papiamento are official in the Caribbean territories.

United Kingdom

United Kingdom

The United Kingdom of Great Britain and Northern Ireland, commonly known as the United Kingdom (UK) or Britain, is a country in Europe, off the north-western coast of the continental mainland. It comprises England, Scotland, Wales and Northern Ireland. The United Kingdom includes the island of Great Britain, the north-eastern part of the island of Ireland, and many smaller islands within the British Isles. Northern Ireland shares a land border with the Republic of Ireland; otherwise, the United Kingdom is surrounded by the Atlantic Ocean, the North Sea, the English Channel, the Celtic Sea and the Irish Sea. The total area of the United Kingdom is 242,495 square kilometres (93,628 sq mi), with an estimated 2023 population of over 68 million people.

New Year's Eve

New Year's Eve

In the Gregorian calendar, New Year's Eve, also known as Old Year's Day or Saint Sylvester's Day in many countries, is the evening or the entire day of the last day of the year, 31 December. The last day of the year is commonly referred to as “New Year’s Eve”. In many countries, New Year's Eve is celebrated with dancing, eating, drinking, and watching or lighting fireworks. Some Christians attend a watchnight service. The celebrations generally go on past midnight into New Year's Day, 1 January.

Restrictions and regulations

United States

The same snails that some people raise or gather as food also are agricultural pests that cause considerable crop damage. Introduced slug and snail varieties tend to be worse pests than native species, probably due in part to the lack of natural controls. Snail pests attack crops ranging from leafy vegetables to fruits that grow near the ground, such as strawberries and tomatoes, to citrus fruits high up on trees.

The Federal Plant Pest Act defines a plant pest as "any living stage (including active and dormant forms) of insects, mites, nematodes, slugs, snails, protozoa, or other invertebrate animals, bacteria, fungi, other parasitic plants or reproductive parts thereof; viruses; or any organisms similar to or allied with any of the foregoing; or any infectious substances, which can directly or indirectly injure or cause disease or damage in or to any plants or parts thereof, or any processed, manufactured, or other products of plants..." The Animal and Plant Health Inspection Service (APHIS) categorizes giant African snails as a "quarantine significant plant pest." The United States does not allow live giant African snails into the country under any circumstances. It is illegal to own or to possess them. APHIS vigorously enforces this regulation and destroys or returns these snails to their country of origin.

Since large infestations of snails can do devastating damage, many states have quarantines against nursery products, and other products, from infested states. Further, it is illegal to import snails (or slugs) into the U.S. without permission from the Plant Protection and Quarantine (PPQ) Division of the Animal and Plant Health Inspection Service, U.S. Department of Agriculture. APHIS also oversees interstate transportation of snails. Anyone who plans to "import, release, or make interstate shipments of" snails, must complete APHIS's PPQ Form 526, Application and Permit to Move Live Plant Pests and Noxious Weeds. This form should be submitted to the state regulatory official. The state will process the request and make a recommendation to APHIS who will then make a decision.

Information on plant pest permits is available at APHIS's website [2].

The Food and Drug Administration (FDA) regulates the canning of low-acid foods such as snails. According to FDA, "establishments engaged in the manufacture of Low-acid or Acidified Canned Foods (LACF) offered for interstate commerce in the United States are required ... to register their facility ... and file scheduled processes for their products with" the FDA. This does not refer to fresh products.[20]

Improper canning of low-acid meats, e.g., snails, involves a risk of botulism. When canning snails for home consumption, carefully follow canning instructions for low-acid meats to prevent food poisoning.

State laws also may apply to imports into certain states and to raising snails in a given state. Some states may want to inspect and approve facilities. Thus, anyone who plans to raise snails also should check with their state's agriculture department.

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Insect

Insect

Insects are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body, three pairs of jointed legs, compound eyes and one pair of antennae. Their blood is not totally contained in vessels; some circulates in an open cavity known as the haemocoel. Insects are the most diverse group of animals; they include more than a million described species and represent more than half of all known living organisms. The total number of extant species is estimated at between six and ten million; potentially over 90% of the animal life forms on Earth are insects. Insects may be found in nearly all environments, although only a small number of species reside in the oceans, which are dominated by another arthropod group, crustaceans, which recent research has indicated insects are nested within.

Mite

Mite

Mites are small arachnids. Mites span two large orders of arachnids, the Acariformes and the Parasitiformes, which were historically grouped together in the subclass Acari, but genetic analysis does not show clear evidence of a close relationship.

Nematode

Nematode

The nematodes or roundworms constitute the phylum Nematoda, with plant-parasitic nematodes also known as eelworms. They are a diverse animal phylum inhabiting a broad range of environments. Less formally, they are categorized as helminths, but are taxonomically classified along with arthropods, tardigrades and other moulting animals in the clade Ecdysozoa, and unlike flatworms, have tubular digestive systems with openings at both ends. Like tardigrades, they have a reduced number of Hox genes, but their sister phylum Nematomorpha has kept the ancestral protostome Hox genotype, which shows that the reduction has occurred within the nematode phylum.

Protozoa

Protozoa

Protozoa are a group of single-celled eukaryotes, either free-living or parasitic, that feed on organic matter such as other microorganisms or organic tissues and debris. Historically, protozoans were regarded as "one-celled animals", because they often possess animal-like behaviours, such as motility and predation, and lack a cell wall, as found in plants and many algae.

Invertebrate

Invertebrate

Invertebrates are a paraphyletic group of animals that neither possess nor develop a vertebral column, derived from the notochord. This is a grouping including all animals apart from the chordate subphylum Vertebrata. Familiar examples of invertebrates include arthropods, mollusks, annelids, echinoderms and cnidarians.

Bacteria

Bacteria

Bacteria are ubiquitous, mostly free-living organisms often consisting of one biological cell. They constitute a large domain of prokaryotic microorganisms. Typically a few micrometres in length, bacteria were among the first life forms to appear on Earth, and are present in most of its habitats. Bacteria inhabit soil, water, acidic hot springs, radioactive waste, and the deep biosphere of Earth's crust. Bacteria are vital in many stages of the nutrient cycle by recycling nutrients such as the fixation of nitrogen from the atmosphere. The nutrient cycle includes the decomposition of dead bodies; bacteria are responsible for the putrefaction stage in this process. In the biological communities surrounding hydrothermal vents and cold seeps, extremophile bacteria provide the nutrients needed to sustain life by converting dissolved compounds, such as hydrogen sulphide and methane, to energy. Bacteria also live in symbiotic and parasitic relationships with plants and animals. Most bacteria have not been characterised and there are many species that cannot be grown in the laboratory. The study of bacteria is known as bacteriology, a branch of microbiology.

Infection

Infection

An infection is the invasion of tissues by pathogens, their multiplication, and the reaction of host tissues to the infectious agent and the toxins they produce. An infectious disease, also known as a transmissible disease or communicable disease, is an illness resulting from an infection.

Animal and Plant Health Inspection Service

Animal and Plant Health Inspection Service

The Animal and Plant Health Inspection Service (APHIS) is an agency of the United States Department of Agriculture (USDA) based in Riverdale, Maryland responsible for protecting animal health, animal welfare, and plant health. APHIS is the lead agency for collaboration with other agencies to protect U.S. agriculture from invasive pests and diseases. APHIS's PPQ is the National Plant Protection Organization for the U.S., and the agency's head of veterinary services/veterinary Deputy Administrator is the Chief Veterinary Officer of the United States.

Plant Protection and Quarantine

Plant Protection and Quarantine

Plant Protection and Quarantine (PPQ) is one of six operational program units within the Animal and Plant Health Inspection Service (APHIS) of the United States Department of Agriculture (USDA). The PPQ works to safeguard agriculture and natural resources in the U.S. against the entry, establishment, and spread of animal and plant pests, and noxious weeds in order to help ensure an abundant, high-quality, and varied food supply.

Food and Drug Administration

Food and Drug Administration

The United States Food and Drug Administration is a federal agency of the Department of Health and Human Services. The FDA is responsible for protecting and promoting public health through the control and supervision of food safety, tobacco products, caffeine products, dietary supplements, prescription and over-the-counter pharmaceutical drugs (medications), vaccines, biopharmaceuticals, blood transfusions, medical devices, electromagnetic radiation emitting devices (ERED), cosmetics, animal foods & feed and veterinary products.

Acid

Acid

An acid is a molecule or ion capable of either donating a proton (i.e. hydrogen ion, H+), known as a Brønsted–Lowry acid, or forming a covalent bond with an electron pair, known as a Lewis acid.

Botulism

Botulism

Botulism is a rare and potentially fatal illness caused by a toxin produced by the bacterium Clostridium botulinum. The disease begins with weakness, blurred vision, feeling tired, and trouble speaking. This may then be followed by weakness of the arms, chest muscles, and legs. Vomiting, swelling of the abdomen, and diarrhea may also occur. The disease does not usually affect consciousness or cause a fever.

Source: "Heliciculture", Wikipedia, Wikimedia Foundation, (2023, February 13th), https://en.wikipedia.org/wiki/Heliciculture.

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References
  1. ^ Gheoca, Voichita (2013-12-01). "Can Heliciculture Act as a Tool for Edible Land Snails' Natural Populations' Management in Romania?" (PDF). Management of Sustainable Development. 5 (2): 21–25. doi:10.2478/msd-2013-0011. ISSN 2247-0220. S2CID 81055812.
  2. ^ Pissia, Maria Α.; Matsakidou, Anthia; Kiosseoglou, Vassilios (2021). "Raw materials from snails for food preparation". Future Foods. 3: 100034. doi:10.1016/j.fufo.2021.100034. S2CID 234819631.
  3. ^ Prehistoric edible land snails in the circum-Mediterranean: the archaeological evidence., D. Lubell. In J-J. Brugal & J. Desse (eds.), Petits Animaux et Sociétés Humaines. Du Complément Alimentaire Aux Ressources Utilitaires. XXIVe rencontres internationales d'archéologie et d'histoire d'Antibes, pp. 77-98. Antibes: Éditions APDCA.
  4. ^ Are land snails a signature for the Mesolithic-Neolithic transition? In, M. Budja (ed.), Neolithic Studies 11. Documenta Praehistorica XXXI: 1-24. D. Lubell.
  5. ^ De Re Rustica III, 12
  6. ^ C. Michael Hogan, Volubilis, The Megalithic Portal, ed. Andy Burnham (2007) [1]
  7. ^ Rebecca Thompson and Sheldon Cheney Raising Snails "National Agricultural Library, Agricultural Research Service, U.S. Department of Agriculture."
  8. ^ Mvodo Meyo, Elise Stephanie; Nkemasong, Zita Akanyi; Shu, Godlove; Ngono, Jean Pierre Nguede; Ngosong, Christopher (2021), Leal Filho, Walter; Pretorius, Rudi; de Sousa, Luiza Olim (eds.), "Snail Farming as an Alternative Profitable Livestock System for Sustainable Development", Sustainable Development in Africa, Cham: Springer International Publishing, pp. 477–490, doi:10.1007/978-3-030-74693-3_26, ISBN 978-3-030-74692-6, S2CID 238016378, retrieved 2022-09-05
  9. ^ "Achatina fulica (giant African land snail)". www.cabi.org. Retrieved 2022-09-05.
  10. ^ "AnimalBase :: Helix pomatia species homepage". www.animalbase.uni-goettingen.de. Retrieved 2022-09-05.
  11. ^ "Snail Farming – Heliciculture". escargot-world.com. Retrieved 2022-09-05.
  12. ^ "European Community Directive on the Conservation of Natural Habitats and of Wild Fauna and Flora (92/43/EEC)" (PDF).
  13. ^ "Edible Molluscs Helicidae". www.manandmollusc.net. Retrieved 2022-09-05.
  14. ^ "AnimalBase :: Cepaea nemoralis species homepage". www.animalbase.uni-goettingen.de. Retrieved 2022-09-05.
  15. ^ "Illegal giant snails threatening America". New York Post. 2014-08-29. Retrieved 2018-03-29.
  16. ^ "1497 : Animal production & health : CTA Publishing". Archived from the original on 2012-10-22.
  17. ^ Bertrand, Jordane (December 15, 2007). "Snail caviar! The new gourmet frontier". Mail & Guardian. Retrieved 21 January 2015.
  18. ^ "Ukraine's snail farmers fear collapse over EU lockdowns". EURACTIV. 27 July 2020. Retrieved 29 July 2020.
  19. ^ a b Willsher, Kim (2021-01-01). "French snail farmers lament sluggish year as Covid crisis dents sales". The Guardian. Retrieved 2021-01-02.
  20. ^ "Acidified and Low-Acid Canned Foods Registration". U.S. Food and Drug Administration. 2022-01-20. Retrieved 2022-03-23.
Further reading
External links
  1. ^ "Archived copy" (PDF). Archived from the original (PDF) on 2006-04-15. Retrieved 2020-01-11.{{cite web}}: CS1 maint: archived copy as title (link)

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