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Freshwater snail

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Bithynia tentaculata, a small freshwater gastropod in the family Bithyniidae
Bithynia tentaculata, a small freshwater gastropod in the family Bithyniidae
Pomacea insularum, an apple snail
Pomacea insularum, an apple snail
Planorbella trivolvis, an air-breathing ramshorn snail
Planorbella trivolvis, an air-breathing ramshorn snail

Freshwater snails are gastropod mollusks that live in fresh water. There are many different families. They are found throughout the world in various habitats, ranging from ephemeral pools to the largest lakes, and from small seeps and springs to major rivers. The great majority of freshwater gastropods have a shell, with very few exceptions. Some groups of snails that live in freshwater respire using gills, whereas other groups need to reach the surface to breathe air. In addition, some are amphibious and have both gills and a lung (e.g. Ampullariidae). Most feed on algae, but many are detritivores and some are filter feeders.

According to a 2008 review of the taxonomy, there are about 4,000 species of freshwater gastropods (3,795–3,972).[1]

At least 33–38 independent lineages of gastropods have successfully colonized freshwater environments.[2] It is not possible to quantify the exact number of these lineages yet, because they have yet to be clarified within the Cerithioidea.[2] From six to eight of these independent lineages occur in North America.[3]

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Fresh water

Fresh water

Fresh water or freshwater is any naturally occurring liquid or frozen water containing low concentrations of dissolved salts and other total dissolved solids. Although the term specifically excludes seawater and brackish water, it does include non-salty mineral-rich waters such as chalybeate springs. Fresh water may encompass frozen and meltwater in ice sheets, ice caps, glaciers, snowfields and icebergs, natural precipitations such as rainfall, snowfall, hail/sleet and graupel, and surface runoffs that form inland bodies of water such as wetlands, ponds, lakes, rivers, streams, as well as groundwater contained in aquifers, subterranean rivers and lakes. Fresh water is the water resource that is of the most and immediate use to humans.

Gastropod shell

Gastropod shell

The gastropod shell is part of the body of a gastropod or snail, a kind of mollusc. The shell is an exoskeleton, which protects from predators, mechanical damage, and dehydration, but also serves for muscle attachment and calcium storage. Some gastropods appear shell-less (slugs) but may have a remnant within the mantle, or in some cases the shell is reduced such that the body cannot be retracted within it (semi-slug). Some snails also possess an operculum that seals the opening of the shell, known as the aperture, which provides further protection. The study of mollusc shells is known as conchology. The biological study of gastropods, and other molluscs in general, is malacology. Shell morphology terms vary by species group.

Pulmonata

Pulmonata

Pulmonata or pulmonates, is an informal group of snails and slugs characterized by the ability to breathe air, by virtue of having a pallial lung instead of a gill, or gills. The group includes many land and freshwater families, and several marine families.

Ampullariidae

Ampullariidae

Ampullariidae, commonly known as the Mystery snails, is a family of large freshwater snails, aquatic gastropod mollusks with a gill and an operculum. These snails simultaneously have a gill and a lung as functional respiratory structures, which are separated by a division of the mantle cavity. This adaptation allows these animals to be amphibious. Species in this family are considered gonochoristic, meaning that each individual organism is either male or female.

Detritivore

Detritivore

Detritivores are heterotrophs that obtain nutrients by consuming detritus. There are many kinds of invertebrates, vertebrates and plants that carry out coprophagy. By doing so, all these detritivores contribute to decomposition and the nutrient cycles. They should be distinguished from other decomposers, such as many species of bacteria, fungi and protists, which are unable to ingest discrete lumps of matter, but instead live by absorbing and metabolizing on a molecular scale. The terms detritivore and decomposer are often used interchangeably, but they describe different organisms. Detritivores are usually arthropods and help in the process of remineralization. Detritivores perform the first stage of remineralization, by fragmenting the dead plant matter, allowing decomposers to perform the second stage of remineralization.

Lineage (evolution)

Lineage (evolution)

An evolutionary lineage is a temporal series of populations, organisms, cells, or genes connected by a continuous line of descent from ancestor to descendant. Lineages are subsets of the evolutionary tree of life. Lineages are often determined by the techniques of molecular systematics.

Cerithioidea

Cerithioidea

The Cerithioidea is a superfamily of marine, brackish water and freshwater gastropod containing more than 200 genera. The Cerithoidea are included unassigned in the subclass Caenogastropoda. The original name of this superfamily was Cerithiacea, in keeping with common superfamily endings at the time.

North America

North America

North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Caribbean Sea, and to the west and south by the Pacific Ocean. Because it is on the North American Tectonic Plate, Greenland is included as a part of North America geographically.

Taxonomy

2005 taxonomy

The following cladogram is an overview of the main clades of gastropods based on the taxonomy of Bouchet & Rocroi (2005),[4] with families that contain freshwater species marked in boldface:[1] (Some of the highlighted families consist entirely of freshwater species, but some of them also contain, or even mainly consist of, marine species.)

† Paleozoic molluscs of uncertain systematic position

† Basal taxa that are certainly Gastropoda

Patellogastropoda

Vetigastropoda

Cocculiniformia

Neritimorpha

† Paleozoic Neritimorpha of uncertain systematic position

Cyrtoneritimorpha

Cycloneritimorpha: Neritiliidae and Neritidae

 Caenogastropoda 

Caenogastropoda of uncertain systematic position

Architaenioglossa: Ampullariidae and Viviparidae

Sorbeoconcha: Melanopsidae, Pachychilidae, Paludomidae, Pleuroceridae, Semisulcospiridae and Thiaridae

 Hypsogastropoda 

Littorinimorpha: Littorinidae, Amnicolidae, Assimineidae, Bithyniidae, Cochliopidae, Helicostoidae, Hydrobiidae, Lithoglyphidae, Moitessieriidae, Pomatiopsidae and Stenothyridae

Ptenoglossa

Neogastropoda: Nassariidae and Marginellidae

Heterobranchia

Lower Heterobranchia: Glacidorbidae and Valvatidae

 Opisthobranchia 

Cephalaspidea

Thecosomata

Gymnosomata

Aplysiomorpha

Acochlidiacea: Acochlidiidae, Tantulidae and Strubelliidae

Sacoglossa

Cylindrobullida

Umbraculida

Nudipleura

Pulmonata

Basommatophora: Chilinidae, Latiidae, Acroloxidae, Lymnaeidae. Planorbidae and Physidae - all these six families together form the clade Hygrophila

Eupulmonata

2010 taxonomy

The following cladogram is an overview of the main clades of gastropods based on the taxonomy of Bouchet & Rocroi (2005),[4] modified after Jörger et al. (2010)[5] and simplified with families that contain freshwater species marked in boldface:[1] (Marine gastropods (Siphonarioidea, Sacoglossa, Amphiboloidea, Pyramidelloidea) are not depicted within Panpulmonata for simplification. Some of these highlighted families consist entirely of freshwater species, but some of them also contain, or even mainly consist of, marine species.)

† Paleozoic molluscs of uncertain systematic position

† Basal taxa that are certainly Gastropoda

Patellogastropoda

Vetigastropoda

Cocculiniformia

Neritimorpha

† Paleozoic Neritimorpha of uncertain systematic position

Cyrtoneritimorpha

Cycloneritimorpha: Neritiliidae and Neritidae

 Caenogastropoda 

Caenogastropoda of uncertain systematic position

Architaenioglossa: Ampullariidae and Viviparidae

Sorbeoconcha: Melanopsidae, Pachychilidae, Paludomidae, Pleuroceridae, Semisulcospiridae and Thiaridae

 Hypsogastropoda 

Littorinimorpha: Littorinidae, Amnicolidae, Assimineidae, Bithyniidae, Cochliopidae, Helicostoidae, Hydrobiidae, Lithoglyphidae, Moitessieriidae, Pomatiopsidae and Stenothyridae

Ptenoglossa

Neogastropoda: Nassariidae and Marginellidae

Heterobranchia

Lower Heterobranchia: Valvatidae

 Euthyneura 

Nudipleura

Euopisthobranchia

Panpulmonata

Glacidorboidea with the only family Glacidorbidae

Hygrophila: Chilinidae, Latiidae, Acroloxidae, Lymnaeidae. Planorbidae and Physidae

Acochlidiacea: Acochlidiidae, Tantulidae and Strubelliidae

Eupulmonata

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Cladogram

Cladogram

A cladogram is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to descendants, nor does it show how much they have changed, so many differing evolutionary trees can be consistent with the same cladogram. A cladogram uses lines that branch off in different directions ending at a clade, a group of organisms with a last common ancestor. There are many shapes of cladograms but they all have lines that branch off from other lines. The lines can be traced back to where they branch off. These branching off points represent a hypothetical ancestor which can be inferred to exhibit the traits shared among the terminal taxa above it. This hypothetical ancestor might then provide clues about the order of evolution of various features, adaptation, and other evolutionary narratives about ancestors. Although traditionally such cladograms were generated largely on the basis of morphological characters, DNA and RNA sequencing data and computational phylogenetics are now very commonly used in the generation of cladograms, either on their own or in combination with morphology.

Patellogastropoda

Patellogastropoda

The Patellogastropoda, common name true limpets and historically called the Docoglossa, are members of a major phylogenetic group of marine gastropods, treated by experts either as a clade or as a taxonomic order.

Neritimorpha

Neritimorpha

Neritimorpha is a taxonomic grouping, an unranked major clade of snails, gastropod mollusks. This grouping includes land snails, sea snails, slugs, some deepwater limpets, and also freshwater snails. Neritimorpha contains around 2,000 extant species. Some Neritimorphs are commonly kept as pets. This clade used to be known as the superorder Neritopsina.

Cyrtoneritimorpha

Cyrtoneritimorpha

Cyrtoneritimorpha, also Cyrtoneritida, is a clade of fossil sea snails, marine gastropod mollusks within the clade Neritimorpha.

Neritiliidae

Neritiliidae

Neritiliidae is a family of submarine cave snails, marine gastropod mollusks or micromollusks in the clade Cycloneritimorpha.

Neritidae

Neritidae

Neritidae, common name the nerites, is a taxonomic family of small to medium-sized saltwater and freshwater snails which have a gill and a distinctive operculum.

Caenogastropoda

Caenogastropoda

Caenogastropoda is a taxonomic subclass of molluscs in the class Gastropoda. It is a large diverse group which are mostly sea snails and other marine gastropod mollusks, but also includes some freshwater snails and some land snails. The subclass is the most diverse and ecologically successful of the gastropods.

Architaenioglossa

Architaenioglossa

Architaenioglossa is a taxonomic group of snails which have gills and often an operculum. They are primarily land and freshwater gastropod mollusks within the clade Caenogastropoda.

Ampullariidae

Ampullariidae

Ampullariidae, commonly known as the Mystery snails, is a family of large freshwater snails, aquatic gastropod mollusks with a gill and an operculum. These snails simultaneously have a gill and a lung as functional respiratory structures, which are separated by a division of the mantle cavity. This adaptation allows these animals to be amphibious. Species in this family are considered gonochoristic, meaning that each individual organism is either male or female.

Melanopsidae

Melanopsidae

Melanopsidae, common name melanopsids, is a family of freshwater gastropods in the clade Sorbeoconcha. Species in this family are native to southern and eastern Europe, northern Africa, parts of the Middle East, New Zealand, and freshwater streams of some large South Pacific islands.

Pachychilidae

Pachychilidae

Pachychilidae, common name pachychilids, is a taxonomic family of freshwater snails, gastropod molluscs in the clade Sorbeoconcha.

Paludomidae

Paludomidae

Paludomidae, common name paludomids, is a family of freshwater snails, gastropod molluscs in the clade Sorbeoconcha.

Neritimorpha

The Neritimorpha are a group of primitive "prosobranch" gilled snails which have a shelly operculum.

  • Neritiliidae, 5 extant freshwater species[1]
  • Neritidae, largely confined to the tropics, also the rivers of Europe, family includes the marine "nerites".[6] There are about 110 extant freshwater species.[1]

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Caenogastropoda

The Caenogastropoda are a large group of gilled operculate snails, which are largely marine. In freshwater habitats there are ten major families of caenogastropods, as well as several other families of lesser importance:

Architaenioglossa
  • Ampullariidae, an exclusively freshwater family that is largely tropical and includes the large "apple snails" kept in aquaria.[6] 105–170 species.[1]
  • Viviparidae, medium to large snails, live-bearing, commonly referred to as "mystery snails". Worldwide except South America, and everywhere confined to fresh waters.[6] 125–150 species.[1]
Sorbeoconcha
  • Melanopsidae, family native to rivers draining to the Mediterranean, also Middle East, and some South Pacific islands.[6] About 25–50 species.[1]
  • Pachychilidae - 165–225 species.[1] native to South and Central America. Formerly included with the Pleuroceridae by many authors.
  • Paludomidae - about 100 species in south Asia, diverse in African Lakes, and Sri Lanka.[1] Formerly classified with the Pleuroceridae by some authors.
  • Pleuroceridae, abundant and diverse in eastern North America, largely high-spired snails of small to large size.[6] About 150 species.[2]
  • Semisulcospiridae, - primarily eastern Asia, Japan, also the Juga snails of northwestern North America. Formerly included with the Pleuroceridae. About 50 species.[2]
  • Thiaridae, high-spired parthenogenic snails of the tropics, includes those referred to as "trumpet snails" in aquaria.[6] About 110 species.[2]
Littorinimorpha
Neogastropoda

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Caenogastropoda

Caenogastropoda

Caenogastropoda is a taxonomic subclass of molluscs in the class Gastropoda. It is a large diverse group which are mostly sea snails and other marine gastropod mollusks, but also includes some freshwater snails and some land snails. The subclass is the most diverse and ecologically successful of the gastropods.

Ampullariidae

Ampullariidae

Ampullariidae, commonly known as the Mystery snails, is a family of large freshwater snails, aquatic gastropod mollusks with a gill and an operculum. These snails simultaneously have a gill and a lung as functional respiratory structures, which are separated by a division of the mantle cavity. This adaptation allows these animals to be amphibious. Species in this family are considered gonochoristic, meaning that each individual organism is either male or female.

Viviparidae

Viviparidae

Viviparidae, sometimes known as the river snails or mystery snails, are a family of large operculate freshwater snails, aquatic gastropod mollusks.

Pomacea bridgesii

Pomacea bridgesii

Pomacea bridgesii, common names the spike-topped apple snail or mystery snail, is a South American species of freshwater snail with gills and an operculum, an aquatic gastropod mollusk in the family Ampullariidae. These snails were most likely introduced to the United States through the aquarium trade.

Melanopsidae

Melanopsidae

Melanopsidae, common name melanopsids, is a family of freshwater gastropods in the clade Sorbeoconcha. Species in this family are native to southern and eastern Europe, northern Africa, parts of the Middle East, New Zealand, and freshwater streams of some large South Pacific islands.

Pachychilidae

Pachychilidae

Pachychilidae, common name pachychilids, is a taxonomic family of freshwater snails, gastropod molluscs in the clade Sorbeoconcha.

Paludomidae

Paludomidae

Paludomidae, common name paludomids, is a family of freshwater snails, gastropod molluscs in the clade Sorbeoconcha.

Pleuroceridae

Pleuroceridae

Pleuroceridae, common name pleurocerids, is a family of small to medium-sized freshwater snails, aquatic gilled gastropod mollusks in the superfamily Cerithioidea.These snails have an operculum and typically a robust high-spired shell.

Semisulcospiridae

Semisulcospiridae

Semisulcospiridae, common name semisulcospirids, is a family of freshwater snails, aquatic gilled gastropod mollusks with an operculum, in the superfamily Cerithioidea.

Thiaridae

Thiaridae

Thiaridae, common name thiarids or trumpet snails, is a family of tropical freshwater snails with an operculum, aquatic gastropod mollusks in the superfamily Cerithioidea.

Melanopsis praemorsa

Melanopsis praemorsa

Melanopsis praemorsa is a species of freshwater snail in the family Melanopsidae.

Semisulcospira kurodai

Semisulcospira kurodai

Semisulcospira kurodai is a species of freshwater snail with an operculum, an aquatic gastropod mollusk in the family Semisulcospiridae. Prior to 2009, this species was classified in the family Pleuroceridae.

Heterobranchia

Acochlidium fijiiensis is one of very few freshwater gastropods without a shell.
Acochlidium fijiiensis is one of very few freshwater gastropods without a shell.
Lower Heterobranchia
Acochlidiacea
Pulmonata, Basommatophora

Basommatophorans are pulmonate or air-breathing aquatic snails, characterized by having their eyes located at the base of their tentacles, rather than at the tips, as in the true land snails Stylommatophora. The majority of basommatophorans have shells that are thin, translucent, and relatively colorless, and all five freshwater basommatophoran families lack an operculum.

  • Chilinidae, small to medium-sized snails confined to temperate and cold South America.[6] About 15 species.[1]
  • Latiidae, small limpet-like snails confined to New Zealand.[6] One[1] or three species.
  • Acroloxidae - about 40 species.[1]
  • Lymnaeidae, found worldwide, but are most numerous in temperate and northern regions.[6] These are the dextral (right-handed) pond snails. About 100 species.
  • Planorbidae, "rams horn" snails, with a worldwide distribution.[6] About 250 species.[1]
  • Physidae, left-handed (sinistral) "pouch snails", native to Europe, Asia, North America.[6] About 80 species.[1]

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Acochlidium fijiiensis

Acochlidium fijiiensis

Acochlidium fijiiensis is a species of freshwater gastropod, an aquatic gastropod mollusc within the family Acochlidiidae. Acochlidium fijiiensis has no shell.

Glacidorbidae

Glacidorbidae

The Glacidorbidae is a taxonomic family of freshwater snails.

Acochlidiacea

Acochlidiacea

Acochlidiacea, common name acochlidians, are a taxonomic clade of very unusual sea snails and sea and freshwater slugs, aquatic gastropod mollusks within the large clade Heterobranchia. Acochlidia is a variant spelling.

Acochlidiidae

Acochlidiidae

Acochlidiidae are a taxonomic family of shell-less freshwater gastropods, aquatic gastropod mollusks within the clade Acochlidiacea.

Acochlidium amboinense

Acochlidium amboinense

Acochlidium amboinense is a species of freshwater gastropod, an aquatic gastropod mollusc within the family Acochlidiidae.

Acochlidium bayerfehlmanni

Acochlidium bayerfehlmanni

Acochlidium bayerfehlmanni is a species of freshwater gastropod, an aquatic gastropod mollusc within the family Acochlidiidae.

Palliohedyle sutteri

Palliohedyle sutteri

Palliohedyle sutteri is a species of gastropods belonging to the family Acochlidiidae.

Strubellia paradoxa

Strubellia paradoxa

Strubellia paradoxa is a species of freshwater slug, a shell-less freshwater gastropod, an aquatic gastropod mollusk within the clade Acochlidiacea.

Basommatophora

Basommatophora

Basommatophora was a term that was previously used as a taxonomic informal group, a group of snails within the informal group Pulmonata, the air-breathing slugs and snails. According to the taxonomy of the Gastropoda, whenever monophyly has not been tested, or where a traditional taxon of gastropods has now been discovered to be paraphyletic or polyphyletic, the term "group" or "informal group" was used.

Stylommatophora

Stylommatophora

Stylommatophora is an order of air-breathing land snails and slugs, terrestrial pulmonate gastropod molluscs. This taxon includes most land snails and slugs.

Acroloxidae

Acroloxidae

Acroloxidae, commonly known as river limpets, are a taxonomic family of very small, freshwater snails, aquatic pulmonate gastropod limpet-like mollusks with a simple flattened conical shell in the clade Hygrophila.

Lymnaeidae

Lymnaeidae

Lymnaeidae, common name the pond snails, is a taxonomic family of small to large air-breathing freshwater snails, aquatic pulmonate gastropod mollusks, that belong to the clade Hygrophila.

Sexual reproduction and self-fertilization

The freshwater snail Physa acuta is in the subclass Heterobranchia and the family Physidae. P. acuta is a self-fertile snail that can undergo either sexual reproduction or self-fertilization. Noel et al.[9] experimentally tested whether accumulation of deleterious mutations is avoided either by inbreeding populations of the snail (undergoing self-fertilization), or in outbreeding populations undergoing sexual reproduction. Inbreeding promotes the homozygous expression of deleterious recessive mutations in progeny that then exposes these mutations to selective elimination because of their deleterious affects on progeny. Outbreeding sexual reproduction allows females to choose male mating partners with smaller mutation loads that then also leads to a reduction of deleterious mutations in progeny. On the basis of their findings, Noel et al.[9] concluded that both outbred and inbred populations of P. acuta can efficiently eliminate deleterious mutations.

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Physella acuta

Physella acuta

Physella acuta is a species of small, left-handed or sinistral, air-breathing freshwater snail, an aquatic gastropod mollusk in the family Physidae. Common names include European physa, tadpole snail, bladder snail, and acute bladder snail. In addition, Physa acuta, Physa heterostropha and Physa integra are synonyms of Physella acuta.

Autogamy

Autogamy

Autogamy, or self-fertilization, refers to the fusion of two gametes that come from one individual. Autogamy is predominantly observed in the form of self-pollination, a reproductive mechanism employed by many flowering plants. However, species of protists have also been observed using autogamy as a means of reproduction. Flowering plants engage in autogamy regularly, while the protists that engage in autogamy only do so in stressful environments.

Mutation

Mutation

In biology, a mutation is an alteration in the nucleic acid sequence of the genome of an organism, virus, or extrachromosomal DNA. Viral genomes contain either DNA or RNA. Mutations result from errors during DNA or viral replication, mitosis, or meiosis or other types of damage to DNA, which then may undergo error-prone repair, cause an error during other forms of repair, or cause an error during replication. Mutations may also result from insertion or deletion of segments of DNA due to mobile genetic elements.

Inbreeding

Inbreeding

Inbreeding is the production of offspring from the mating or breeding of individuals or organisms that are closely related genetically. By analogy, the term is used in human reproduction, but more commonly refers to the genetic disorders and other consequences that may arise from expression of deleterious or recessive traits resulting from incestuous sexual relationships and consanguinity. Animals avoid incest only rarely.

Outcrossing

Outcrossing

Out-crossing or out-breeding is the technique of crossing between different breeds. This is the practice of introducing distantly related genetic material into a breeding line, thereby increasing genetic diversity.

Sexual reproduction

Sexual reproduction

Sexual reproduction is a type of reproduction that involves a complex life cycle in which a gamete with a single set of chromosomes combines with another gamete to produce a zygote that develops into an organism composed of cells with two sets of chromosomes (diploid). This is typical in animals, though the number of chromosome sets and how that number changes in sexual reproduction varies, especially among plants, fungi, and other eukaryotes.

Zygosity

Zygosity

Zygosity is the degree to which both copies of a chromosome or gene have the same genetic sequence. In other words, it is the degree of similarity of the alleles in an organism.

As human food

Several different freshwater snail species are eaten in Asian cuisine.

Archaeological investigations in Guatemala have revealed that the diet of the Maya of the Classic Period (AD 250–900) included freshwater snails.[10]

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Asian cuisine

Asian cuisine

Asian food incorporates a few significant provincial cooking styles: Central Asian, East Asian, North Asian, South Asian, Southeast Asian, and West Asian. A food is a trademark way of cooking practices and customs, usually associated with a specific culture. Asia, being the largest and most populous continent, is home to many cultures, many of which have their own characteristic cuisine. Asian cuisine are also famous about their spices, Asian people traditionally use different kind of spices in their regular meals.

Guatemala

Guatemala

Guatemala, officially the Republic of Guatemala, is a country in Central America. It is bordered to the north and west by Mexico, to the northeast by Belize and the Caribbean, to the east by Honduras, to the southeast by El Salvador and to the south by the Pacific Ocean. With an estimated population of around 17.6 million, Guatemala is the most populous country in Central America and the 11th most populous country in the Americas. It is a representative democracy with its capital and largest city being Nueva Guatemala de la Asunción, also known as Guatemala City, the most populous city in Central America.

Maya civilization

Maya civilization

The Maya civilization of the Mesoamerican people is known by its ancient temples and glyphs. Its Maya script is the most sophisticated and highly developed writing system in the pre-Columbian Americas. It is also noted for its art, architecture, mathematics, calendar, and astronomical system.

Mesoamerican chronology

Mesoamerican chronology

Mesoamerican chronology divides the history of prehispanic Mesoamerica into several periods: the Paleo-Indian ; the Archaic, the Preclassic or Formative (2500 BCE – 250 CE), the Classic (250–900 CE), and the Postclassic (900–1521 CE); as well as the post European contact Colonial Period (1521–1821), and Postcolonial, or the period after independence from Spain (1821–present).

Ampullariidae

Ampullariidae

Ampullariidae, commonly known as the Mystery snails, is a family of large freshwater snails, aquatic gastropod mollusks with a gill and an operculum. These snails simultaneously have a gill and a lung as functional respiratory structures, which are separated by a division of the mantle cavity. This adaptation allows these animals to be amphibious. Species in this family are considered gonochoristic, meaning that each individual organism is either male or female.

Viviparidae

Viviparidae

Viviparidae, sometimes known as the river snails or mystery snails, are a family of large operculate freshwater snails, aquatic gastropod mollusks.

Poipet

Poipet

Poipet is a boomtown on the Cambodian-Thai border, in Poipet Municipality, Banteay Meanchey Province. It is a key crossing point between the two countries, and also extremely popular as a gambling destination as gambling is popular, but mostly illegal in Thailand.

Cambodia

Cambodia

Cambodia, officially the Kingdom of Cambodia, is a country located in the southern portion of the Indochinese Peninsula in Southeast Asia, spanning an area of 181,035 square kilometres, bordered by Thailand to the northwest, Laos to the north, Vietnam to the east, and the Gulf of Thailand to the southwest. The capital and largest city is Phnom Penh.

Kolkata

Kolkata

Kolkata is the capital of the Indian state of West Bengal, on the eastern bank of the Hooghly River 80 km (50 mi) west of the border with Bangladesh. It is the primary business, commercial, and financial hub of Eastern India and the main port of communication for North-East India. According to the 2011 Indian census, Kolkata is the seventh-most populous city in India, with a population of 45 lakh (4.5 million) residents within the city. It is a part of Kolkata Metropolitan Area which has a population of over 1.41 crore (14.1 million) residents making it the third-most populous metropolitan area in India. In 2021, the Kolkata metropolitan area crossed 1.5 crore (15 million) registered voters. The Port of Kolkata is India's oldest operating port and its sole major riverine port. Kolkata is regarded as the cultural capital of India. Kolkata is the second largest Bengali-speaking city after Dhaka. It has the highest number of nobel laureates among all cities in India.

West Bengal

West Bengal

West Bengal is a state in the eastern portion of India. It is situated along the Bay of Bengal, along with a population of over 91 million inhabitants within an area of 88,752 km2 (34,267 sq mi). West Bengal is the fourth-most populous and thirteenth-largest state by area in India, as well as the eighth-most populous country subdivision of the world. As a part of the Bengal region of the Indian subcontinent, it borders Bangladesh in the east, and Nepal and Bhutan in the north. It also borders the Indian states of Odisha, Jharkhand, Bihar, Sikkim and Assam. The state capital is Kolkata, the third-largest metropolis, and seventh largest city by population in India. West Bengal includes the Darjeeling Himalayan hill region, the Ganges delta, the Rarh region, the coastal Sundarbans and the Bay of Bengal. The state's main ethnic group are the Bengalis, with the Bengali Hindus forming the demographic majority.

India

India

India, officially the Republic of India, is a country in South Asia. It is the seventh-largest country by area and the second-most populous country. Bounded by the Indian Ocean on the south, the Arabian Sea on the southwest, and the Bay of Bengal on the southeast, it shares land borders with Pakistan to the west; China, Nepal, and Bhutan to the north; and Bangladesh and Myanmar to the east. In the Indian Ocean, India is in the vicinity of Sri Lanka and the Maldives; its Andaman and Nicobar Islands share a maritime border with Thailand, Myanmar, and Indonesia.

Aquarium snails

Freshwater snails are commonly found in aquaria along with tropical fish. Species available vary in different parts of the world. In the United States, commonly available species include ramshorn snails such as Planorbella duryi, apple snails such as Pomacea bridgesii, the high-spired thiarid Malaysian trumpet snail, Melanoides tuberculata, and several Neritina species.

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Parasitology

Life cycle of two liver fluke species which have freshwater snails as intermediate hosts
Life cycle of two liver fluke species which have freshwater snails as intermediate hosts

Freshwater snails are widely known to be hosts in the lifecycles of a variety of human and animal parasites, particularly trematodes or "flukes". Some of these relations for prosobranch snails include Oncomelania in the family Pomatiopsidae as hosts of Schistosoma, and Bithynia, Parafossarulus and Amnicola as hosts of Opisthorchis.[11] Thiara and Semisulcospira may host Paragonimus.[11] Juga plicifera may host Nanophyetus salmincola.[12] Basommatophoran snails are even more widely infected, with many Biomphalaria (Planorbidae) serving as hosts for Schistosoma mansoni, Fasciolopsis and other parasitic groups.[11] The tiny Bulinus snails are hosts for Schistosoma haematobium.[11] Lymnaeid snails (Lymnaeidae) serve as hosts for Fasciola and the cerceriae causing swimmer's itch.[11] The term "neglected tropical diseases" applies to all snail-borne infections, including schistosomiasis, fascioliasis, fasciolopsiasis, paragonimiasis, opisthorchiasis, clonorchiasis, and angiostrongyliasis.[13]

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Liver fluke

Liver fluke

Liver fluke is a collective name of a polyphyletic group of parasitic trematodes under the phylum Platyhelminthes. They are principally parasites of the liver of various mammals, including humans. Capable of moving along the blood circulation, they can occur also in bile ducts, gallbladder, and liver parenchyma. In these organs, they produce pathological lesions leading to parasitic diseases. They have complex life cycles requiring two or three different hosts, with free-living larval stages in water.

Parasitism

Parasitism

Parasitism is a close relationship between species, where one organism, the parasite, lives on or inside another organism, the host, causing it some harm, and is adapted structurally to this way of life. The entomologist E. O. Wilson has characterised parasites as "predators that eat prey in units of less than one". Parasites include single-celled protozoans such as the agents of malaria, sleeping sickness, and amoebic dysentery; animals such as hookworms, lice, mosquitoes, and vampire bats; fungi such as honey fungus and the agents of ringworm; and plants such as mistletoe, dodder, and the broomrapes.

Oncomelania

Oncomelania

Oncomelania is a genus of very small tropical freshwater snails, aquatic gastropod mollusks in the family Pomatiopsidae.

Bithynia (gastropod)

Bithynia (gastropod)

Bithynia is a genus of small freshwater snails with an operculum, aquatic prosobranch gastropod mollusks in the family Bithyniidae.

Parafossarulus

Parafossarulus

Parafossarulus is a genus of freshwater snails with gills and an operculum, an aquatic prosobranch gastropod mollusks in the family Bithyniidae.

Amnicola

Amnicola

Amnicola is a genus of very small freshwater snails which have an operculum. Amnicola species are aquatic prosobranch gastropod mollusks in the family Amnicolidae according to the taxonomy of the Gastropoda.

Opisthorchis

Opisthorchis

Opisthorchis is a genus of flukes in the family Opisthorchiidae.

Paragonimus

Paragonimus

Paragonimus is a genus of flukes (trematodes) and is the only genus in the monotypic family Paragonimidae. Some tens of species have been described, but they are difficult to distinguish, so it is not clear how many of the named species may be synonyms. The name Paragonimus is derived from the combination of two Greek words, “para” and “gonimos”. Several of the species are known as lung flukes. In humans some of the species occur as zoonoses; the term for the condition is paragonimiasis. The first intermediate hosts of Paragonimus include at least 54 species of freshwater snails from superfamilies Cerithioidea and Rissooidea.

Juga plicifera

Juga plicifera

Juga plicifera, common name pleated juga, and graceful keeled horn snail is a species of small freshwater snail with an ovate and corneous operculum, ranging in size from 16mm to 35mm long. The snail is dextrally coiled with about 15 whorls and 10 to 12 axial plicae on each whorl. It is an aquatic gastropod mollusk in the family Semisulcospiridae.

Biomphalaria

Biomphalaria

Biomphalaria is a genus of air-breathing freshwater snails, aquatic pulmonate gastropod mollusks in the family Planorbidae, the ram's horn snails and their allies.

Fasciolopsis

Fasciolopsis

Fasciolopsis is a genus of trematodes. They are also known as giant intestinal flukes.

Bulinus

Bulinus

Bulinus is a genus of small tropical freshwater snails, aquatic gastropod mollusks in the family Bulinidae, the ramshorn snails and their allies.

Source: "Freshwater snail", Wikipedia, Wikimedia Foundation, (2023, March 15th), https://en.wikipedia.org/wiki/Freshwater_snail.

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References

This article incorporates CC-BY-2.5 text from the reference[13]

  1. ^ a b c d e f g h i j k l m n o p q r s t u v w x y z aa ab ac ad ae Strong E. E., Gargominy O., Ponder W. F. & Bouchet P. (2008). "Global Diversity of Gastropods (Gastropoda; Mollusca) in Freshwater". Hydrobiologia 595: 149–166. hdl.handle.net doi:10.1007/s10750-007-9012-6.
  2. ^ a b c d e Strong E. E., Colgan D. J., Healy J. M., Lydeard C., Ponder W. F. & Glaubrecht M. (2011). "Phylogeny of the gastropod superfamily Cerithioidea using morphology and molecules". Zoological Journal of the Linnean Society 162(1): 43–89. doi:10.1111/j.1096-3642.2010.00670.x.
  3. ^ Dillon R. T. (2006). Chapter 21. Freshwater Gastropoda. pages 251–259. In: Sturm C. F., Pearce T. A. & Valdés A. (eds.) (2006). The Mollusks: A Guide to Their Study, Collection, and Preservation. American Malacological Society, 445 pp. ISBN 978-1-58112-930-4.
  4. ^ a b Bouchet, Philippe; Rocroi, Jean-Pierre; Frýda, Jiri; Hausdorf, Bernard; Ponder, Winston; Valdés, Ángel & Warén, Anders (2005). "Classification and nomenclator of gastropod families". Malacologia. Hackenheim, Germany: ConchBooks. 47 (1–2): 1–397. ISBN 3-925919-72-4. ISSN 0076-2997.
  5. ^ Jörger K. M., Stöger I., Kano Y., Fukuda H., Knebelsberger T. & Schrödl M. (2010). "On the origin of Acochlidia and other enigmatic euthyneuran gastropods, with implications for the systematics of Heterobranchia". BMC Evolutionary Biology 10: 323. doi:10.1186/1471-2148-10-323.
  6. ^ a b c d e f g h i j k l m n Banarescu P. (1990). Zoogeography of Fresh Waters, Vol. 1, General Distribution and Dispersal of Freshwater Animals. AULA - Verlag, Weisbaden.
  7. ^ Reid D. G., Aravind N. A., & Madhyastha N. A. (2013). "A unique radiation of marine littorinid snails in the freshwater streams of the Western Ghats of India: the genus Cremnoconchus W.T. Blanford, 1869 (Gastropoda: Littorinidae)". Zoological Journal of the Linnean Society 167(1): 93–135. doi:10.1111/j.1096-3642.2012.00875.x.
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  9. ^ a b Noël E, Fruitet E, Lelaurin D, Bonel N, Ségard A, Sarda V, Jarne P, David P. Sexual selection and inbreeding: Two efficient ways to limit the accumulation of deleterious mutations. Evol Lett. 2018 Dec 10;3(1):80-92. doi: 10.1002/evl3.93. PMID: 30788144; PMCID: PMC6369961
  10. ^ Foias A. E. (2000). "Entre la política y economía: Resultados preliminares de las primeras temporadas del Proyecto Arqueológico Motul de San José" (PDF). XIII Simposio de Investigaciones Arqueológicas en Guatemala, 1999 (Edited by J.P. Laporte, H. Escobedo, B. Arroyo and A.C. De Suasnávar) (in Spanish): 771–799. Archived from the original (PDF online publication) on 2009-03-18. Retrieved 2009-03-01., page 777.
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Further reading

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