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Animal

From Wikipedia, in a visual modern way

Animals are multicellular, eukaryotic organisms in the biological kingdom Animalia. With few exceptions, animals consume organic material, breathe oxygen, are able to move, can reproduce sexually, and grow from a hollow sphere of cells, the blastula, during embryonic development. Over 1.5 million living animal species have been described—of which around 1 million are insects—but it has been estimated there are over 7 million animal species in total. Animals range in length from 8.5 micrometres (0.00033 in) to 33.6 metres (110 ft). They have complex interactions with each other and their environments, forming intricate food webs. The scientific study of animals is known as zoology.

Most living animal species are in Bilateria, a clade whose members have a bilaterally symmetric body plan. The Bilateria include the protostomes, containing animals such as nematodes, arthropods, flatworms, annelids and molluscs, and the deuterostomes, containing the echinoderms and the chordates, the latter including the vertebrates. Life forms interpreted as early animals were present in the Ediacaran biota of the late Precambrian. Many modern animal phyla became clearly established in the fossil record as marine species during the Cambrian explosion, which began around 539 million years ago. 6,331 groups of genes common to all living animals have been identified; these may have arisen from a single common ancestor that lived 650 million years ago.

Historically, Aristotle divided animals into those with blood and those without. Carl Linnaeus created the first hierarchical biological classification for animals in 1758 with his Systema Naturae, which Jean-Baptiste Lamarck expanded into 14 phyla by 1809. In 1874, Ernst Haeckel divided the animal kingdom into the multicellular Metazoa (now synonymous with Animalia) and the Protozoa, single-celled organisms no longer considered animals. In modern times, the biological classification of animals relies on advanced techniques, such as molecular phylogenetics, which are effective at demonstrating the evolutionary relationships between taxa.

Humans make use of many animal species, such as for food (including meat, milk, and eggs), for materials (such as leather and wool), as pets, and as working animals including for transport. Dogs have been used in hunting, as have birds of prey, while many terrestrial and aquatic animals were hunted for sports. Nonhuman animals have appeared in art from the earliest times and are featured in mythology and religion.

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Bilateria

Bilateria

Bilateria is a group of animals, called bilaterians, with bilateral symmetry as an embryo. This also means they have a head and a tail, as well as a belly and a back. Nearly all are bilaterally symmetrical as adults as well; the most notable exception is the echinoderms, which achieve secondary pentaradial symmetry as adults, but are bilaterally symmetrical during embryonic development.

Clade

Clade

In biological phylogenetics, a clade, also known as a monophyletic group or natural group, is a grouping of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. In the taxonomical literature, sometimes the Latin form cladus is used rather than the English form.

Body plan

Body plan

A body plan, Bauplan, or ground plan is a set of morphological features common to many members of a phylum of animals. The vertebrates share one body plan, while invertebrates have many.

Arthropod

Arthropod

Arthropods are invertebrate animals with an exoskeleton, a segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and cuticle made of chitin, often mineralised with calcium carbonate. The arthropod body plan consists of segments, each with a pair of appendages. Arthropods are bilaterally symmetrical and their body possesses an external skeleton. In order to keep growing, they must go through stages of moulting, a process by which they shed their exoskeleton to reveal a new one. Some species have wings. They are an extremely diverse group, with up to 10 million species.

Annelid

Annelid

The annelids, also known as the segmented worms, are a large phylum, with over 22,000 extant species including ragworms, earthworms, and leeches. The species exist in and have adapted to various ecologies – some in marine environments as distinct as tidal zones and hydrothermal vents, others in fresh water, and yet others in moist terrestrial environments.

Chordate

Chordate

A chordate is an animal of the phylum Chordata. All chordates possess, at some point during their larval or adult stages, five synapomorphies, or primary physical characteristics, that distinguish them from all the other taxa. These five synapomorphies include a notochord, dorsal hollow nerve cord, endostyle or thyroid, pharyngeal slits, and a post-anal tail. The name “chordate” comes from the first of these synapomorphies, the notochord, which plays a significant role in chordate structure and movement. Chordates are also bilaterally symmetric, have a coelom, possess a circulatory system, and exhibit metameric segmentation.

Cambrian explosion

Cambrian explosion

The Cambrian explosion, Cambrian radiation, Cambrian diversification, or the Biological Big Bang refers to an interval of time approximately 538.8 million years ago in the Cambrian Period when practically all major animal phyla started appearing in the fossil record. It lasted for about 13 – 25 million years and resulted in the divergence of most modern metazoan phyla. The event was accompanied by major diversification in other groups of organisms as well.

Cryogenian

Cryogenian

The Cryogenian is a geologic period that lasted from 720 to 635 million years ago. It forms the second geologic period of the Neoproterozoic Era, preceded by the Tonian Period and followed by the Ediacaran.

Aristotle's biology

Aristotle's biology

Aristotle's biology is the theory of biology, grounded in systematic observation and collection of data, mainly zoological, embodied in Aristotle's books on the science. Many of his observations were made during his stay on the island of Lesbos, including especially his descriptions of the marine biology of the Pyrrha lagoon, now the Gulf of Kalloni. His theory is based on his concept of form, which derives from but is markedly unlike Plato's theory of Forms.

Carl Linnaeus

Carl Linnaeus

Carl Linnaeus, also known after his ennoblement in 1761 as Carl von Linné, was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the modern system of naming organisms. He is known as the "father of modern taxonomy". Many of his writings were in Latin; his name is rendered in Latin as Carolus Linnæus and, after his 1761 ennoblement, as Carolus a Linné.

Dog

Dog

The dog is a domesticated descendant of the wolf. Also called the domestic dog, it is derived from the extinct Pleistocene wolf, and the modern wolf is the dog's nearest living relative. Dogs were the first species to be domesticated by hunter-gatherers over 15,000 years ago before the development of agriculture. Due to their long association with humans, dogs have expanded to a large number of domestic individuals and gained the ability to thrive on a starch-rich diet that would be inadequate for other canids.

Aquatic animal

Aquatic animal

An aquatic animal is any animal, whether vertebrate or invertebrate, that lives in water for all or most of its lifetime. Many insects such as mosquitoes, mayflies, dragonflies and caddisflies have aquatic larvae, with winged adults. Aquatic animals may breathe air or extract oxygen from water through specialised organs called gills, or directly through the skin. Natural environments and the animals that live in them can be categorized as aquatic (water) or terrestrial (land). This designation is polyphyletic.

Etymology

The word "animal" comes from the Latin animalis, meaning 'having breath', 'having soul' or 'living being'.[8] The biological definition includes all members of the kingdom Animalia.[9] In colloquial usage, the term animal is often used to refer only to nonhuman animals.[10][11][12][13] The term "metazoa" is derived from the Ancient Greek μετα (meta, meaning "later") and ζῷᾰ (zōia, plural of ζῷον zōion, meaning animal).[14][15]

Characteristics

Animals are unique in having the ball of cells of the early embryo (1) develop into a hollow ball or blastula (2).
Animals are unique in having the ball of cells of the early embryo (1) develop into a hollow ball or blastula (2).

Animals have several characteristics that set them apart from other living things. Animals are eukaryotic and multicellular.[16][17] Unlike plants and algae, which produce their own nutrients,[18] animals are heterotrophic,[17][19] feeding on organic material and digesting it internally.[20] With very few exceptions, animals respire aerobically.[a][22] All animals are motile[23] (able to spontaneously move their bodies) during at least part of their life cycle, but some animals, such as sponges, corals, mussels, and barnacles, later become sessile. The blastula is a stage in embryonic development that is unique to animals, allowing cells to be differentiated into specialised tissues and organs.[24]

Structure

All animals are composed of cells, surrounded by a characteristic extracellular matrix composed of collagen and elastic glycoproteins.[25] During development, the animal extracellular matrix forms a relatively flexible framework upon which cells can move about and be reorganised, making the formation of complex structures possible. This may be calcified, forming structures such as shells, bones, and spicules.[26] In contrast, the cells of other multicellular organisms (primarily algae, plants, and fungi) are held in place by cell walls, and so develop by progressive growth.[27] Animal cells uniquely possess the cell junctions called tight junctions, gap junctions, and desmosomes.[28]

With few exceptions—in particular, the sponges and placozoans—animal bodies are differentiated into tissues.[29] These include muscles, which enable locomotion, and nerve tissues, which transmit signals and coordinate the body. Typically, there is also an internal digestive chamber with either one opening (in Ctenophora, Cnidaria, and flatworms) or two openings (in most bilaterians).[30]

Reproduction and development

Sexual reproduction is nearly universal in animals, such as these dragonflies.
Sexual reproduction is nearly universal in animals, such as these dragonflies.

Nearly all animals make use of some form of sexual reproduction.[31] They produce haploid gametes by meiosis; the smaller, motile gametes are spermatozoa and the larger, non-motile gametes are ova.[32] These fuse to form zygotes,[33] which develop via mitosis into a hollow sphere, called a blastula. In sponges, blastula larvae swim to a new location, attach to the seabed, and develop into a new sponge.[34] In most other groups, the blastula undergoes more complicated rearrangement.[35] It first invaginates to form a gastrula with a digestive chamber and two separate germ layers, an external ectoderm and an internal endoderm.[36] In most cases, a third germ layer, the mesoderm, also develops between them.[37] These germ layers then differentiate to form tissues and organs.[38]

Repeated instances of mating with a close relative during sexual reproduction generally leads to inbreeding depression within a population due to the increased prevalence of harmful recessive traits.[39][40] Animals have evolved numerous mechanisms for avoiding close inbreeding.[41]

Some animals are capable of asexual reproduction, which often results in a genetic clone of the parent. This may take place through fragmentation; budding, such as in Hydra and other cnidarians; or parthenogenesis, where fertile eggs are produced without mating, such as in aphids.[42][43]

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Embryo

Embryo

An embryo is an initial stage of development of a multicellular organism. In organisms that reproduce sexually, embryonic development is the part of the life cycle that begins just after fertilization of the female egg cell by the male sperm cell. The resulting fusion of these two cells produces a single-celled zygote that undergoes many cell divisions that produce cells known as blastomeres. The blastomeres are arranged as a solid ball that when reaching a certain size, called a morula, takes in fluid to create a cavity called a blastocoel. The structure is then termed a blastula, or a blastocyst in mammals.

Autotroph

Autotroph

An autotroph is an organism that produces complex organic compounds using carbon from simple substances such as carbon dioxide, generally using energy from light (photosynthesis) or inorganic chemical reactions (chemosynthesis). They convert an abiotic source of energy into energy stored in organic compounds, which can be used by other organisms. Autotrophs do not need a living source of carbon or energy and are the producers in a food chain, such as plants on land or algae in water. Autotrophs can reduce carbon dioxide to make organic compounds for biosynthesis and as stored chemical fuel. Most autotrophs use water as the reducing agent, but some can use other hydrogen compounds such as hydrogen sulfide.

Heterotroph

Heterotroph

A heterotroph is an organism that cannot produce its own food, instead taking nutrition from other sources of organic carbon, mainly plant or animal matter. In the food chain, heterotrophs are primary, secondary and tertiary consumers, but not producers. Living organisms that are heterotrophic include all animals and fungi, some bacteria and protists, and many parasitic plants. The term heterotroph arose in microbiology in 1946 as part of a classification of microorganisms based on their type of nutrition. The term is now used in many fields, such as ecology in describing the food chain.

Motility

Motility

Motility is the ability of an organism to move independently, using metabolic energy.

Biological life cycle

Biological life cycle

In biology, a biological life cycle is a series of changes in form that an organism undergoes, returning to the starting state. "The concept is closely related to those of the life history, development and ontogeny, but differs from them in stressing renewal." Transitions of form may involve growth, asexual reproduction, or sexual reproduction.

Coral

Coral

Corals are marine invertebrates within the class Anthozoa of the phylum Cnidaria. They typically form compact colonies of many identical individual polyps. Coral species include the important reef builders that inhabit tropical oceans and secrete calcium carbonate to form a hard skeleton.

Mussel

Mussel

Mussel is the common name used for members of several families of bivalve molluscs, from saltwater and freshwater habitats. These groups have in common a shell whose outline is elongated and asymmetrical compared with other edible clams, which are often more or less rounded or oval.

Barnacle

Barnacle

Barnacles are a type of arthropod constituting the subclass Cirripedia in the subphylum Crustacea, and are hence related to crabs and lobsters. Barnacles are exclusively marine, and tend to live in shallow and tidal waters, typically in erosive settings. They are sessile (nonmobile) and most are suspension feeders, but those in infraclass Rhizocephala are highly specialized parasites on crustaceans. They have four nektonic larval stages. Around 1,000 barnacle species are currently known. The name Cirripedia is Latin, meaning "curl-footed". The study of barnacles is called cirripedology.

Sessility (motility)

Sessility (motility)

Sessility is the biological property of an organism describing its lack of a means of self-locomotion. Sessile organisms for which natural motility is absent are normally immobile. This is distinct from the botanical concept of sessility, which refers to an organism or biological structure attached directly by its base without a stalk.

Cellular differentiation

Cellular differentiation

Cellular differentiation is the process in which a stem cell changes from one type to a differentiated one. Usually, the cell changes to a more specialized type. Differentiation happens multiple times during the development of a multicellular organism as it changes from a simple zygote to a complex system of tissues and cell types. Differentiation continues in adulthood as adult stem cells divide and create fully differentiated daughter cells during tissue repair and during normal cell turnover. Some differentiation occurs in response to antigen exposure. Differentiation dramatically changes a cell's size, shape, membrane potential, metabolic activity, and responsiveness to signals. These changes are largely due to highly controlled modifications in gene expression and are the study of epigenetics. With a few exceptions, cellular differentiation almost never involves a change in the DNA sequence itself. However, metabolic composition does get altered quite dramatically where stem cells are characterized by abundant metabolites with highly unsaturated structures whose levels decrease upon differentiation. Thus, different cells can have very different physical characteristics despite having the same genome.

Extracellular matrix

Extracellular matrix

In biology, the extracellular matrix (ECM), also called intercellular matrix, is a three-dimensional network consisting of extracellular macromolecules and minerals, such as collagen, enzymes, glycoproteins and hydroxyapatite that provide structural and biochemical support to surrounding cells. Because multicellularity evolved independently in different multicellular lineages, the composition of ECM varies between multicellular structures; however, cell adhesion, cell-to-cell communication and differentiation are common functions of the ECM.

Collagen

Collagen

Collagen is the main structural protein in the extracellular matrix found in the body's various connective tissues. As the main component of connective tissue, it is the most abundant protein in mammals, making up from 25% to 35% of the whole-body protein content. Collagen consists of amino acids bound together to form a triple helix of elongated fibril known as a collagen helix. It is mostly found in connective tissue such as cartilage, bones, tendons, ligaments, and skin.

Ecology

Predators, such as this ultramarine flycatcher (Ficedula superciliaris), feed on other animals.
Predators, such as this ultramarine flycatcher (Ficedula superciliaris), feed on other animals.

Animals are categorised into ecological groups depending on how they obtain or consume organic material, including carnivores, herbivores, omnivores, detritivores,[44] and parasites.[45] Interactions between animals form complex food webs. In carnivorous or omnivorous species, predation is a consumer–resource interaction where a predator feeds on another organism (called its prey).[46] Selective pressures imposed on one another lead to an evolutionary arms race between predator and prey, resulting in various anti-predator adaptations.[47][48] Almost all multicellular predators are animals.[49] Some consumers use multiple methods; for example, in parasitoid wasps, the larvae feed on the hosts' living tissues, killing them in the process,[50] but the adults primarily consume nectar from flowers.[51] Other animals may have very specific feeding behaviours, such as hawksbill sea turtles primarily eating sponges.[52]

Hydrothermal vent mussels and shrimps
Hydrothermal vent mussels and shrimps

Most animals rely on the biomass and energy produced by plants through photosynthesis. Herbivores eat plant material directly, while carnivores, and other animals on higher trophic levels typically acquire it indirectly by eating other animals. Animals oxidize carbohydrates, lipids, proteins, and other biomolecules, which allows the animal to grow and to sustain biological processes such as locomotion.[53][54][55] Animals living close to hydrothermal vents and cold seeps on the dark sea floor consume organic matter of archaea and bacteria produced in these locations through chemosynthesis (by oxidizing inorganic compounds, such as hydrogen sulfide).[56]

Animals originally evolved in the sea. Lineages of arthropods colonised land around the same time as land plants, probably between 510 and 471 million years ago during the Late Cambrian or Early Ordovician.[57] Vertebrates such as the lobe-finned fish Tiktaalik started to move on to land in the late Devonian, about 375 million years ago.[58][59] Animals occupy virtually all of earth's habitats and microhabitats, including salt water, hydrothermal vents, fresh water, hot springs, swamps, forests, pastures, deserts, air, and the interiors of animals, plants, fungi and rocks.[60] Animals are however not particularly heat tolerant; very few of them can survive at constant temperatures above 50 °C (122 °F).[61] Only very few species of animals (mostly nematodes) inhabit the most extreme cold deserts of continental Antarctica.[62]

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Ecology

Ecology

Ecology is the study of the relationships among living organisms, including humans, and their physical environment. Ecology considers organisms at the individual, population, community, ecosystem, and biosphere level. Ecology overlaps with the closely related sciences of biogeography, evolutionary biology, genetics, ethology, and natural history. Ecology is a branch of biology, and it is not synonymous with environmentalism.

Carnivore

Carnivore

A carnivore, or meat-eater, is an animal or plant whose food and energy requirements derive from animal tissues whether through hunting or scavenging.

Herbivore

Herbivore

A herbivore is an animal anatomically and physiologically adapted to eating plant material, for example foliage or marine algae, for the main component of its diet. As a result of their plant diet, herbivorous animals typically have mouthparts adapted to rasping or grinding. Horses and other herbivores have wide flat teeth that are adapted to grinding grass, tree bark, and other tough plant material.

Omnivore

Omnivore

An omnivore is an animal that has the ability to eat and survive on both plant and animal matter. Obtaining energy and nutrients from plant and animal matter, omnivores digest carbohydrates, protein, fat, and fiber, and metabolize the nutrients and energy of the sources absorbed. Often, they have the ability to incorporate food sources such as algae, fungi, and bacteria into their diet.

Detritivore

Detritivore

Detritivores are heterotrophs that obtain nutrients by consuming detritus. There are many kinds of invertebrates, vertebrates and plants that carry out coprophagy. By doing so, all these detritivores contribute to decomposition and the nutrient cycles. They should be distinguished from other decomposers, such as many species of bacteria, fungi and protists, which are unable to ingest discrete lumps of matter, but instead live by absorbing and metabolizing on a molecular scale. The terms detritivore and decomposer are often used interchangeably, but they describe different organisms. Detritivores are usually arthropods and help in the process of remineralization. Detritivores perform the first stage of remineralization, by fragmenting the dead plant matter, allowing decomposers to perform the second stage of remineralization.

Food web

Food web

A food web is the natural interconnection of food chains and a graphical representation of what-eats-what in an ecological community. Another name for food web is consumer-resource system. Ecologists can broadly lump all life forms into one of two categories called trophic levels: 1) the autotrophs, and 2) the heterotrophs. To maintain their bodies, grow, develop, and to reproduce, autotrophs produce organic matter from inorganic substances, including both minerals and gases such as carbon dioxide. These chemical reactions require energy, which mainly comes from the Sun and largely by photosynthesis, although a very small amount comes from bioelectrogenesis in wetlands, and mineral electron donors in hydrothermal vents and hot springs. These trophic levels are not binary, but form a gradient that includes complete autotrophs, which obtain their sole source of carbon from the atmosphere, mixotrophs, which are autotrophic organisms that partially obtain organic matter from sources other than the atmosphere, and complete heterotrophs that must feed to obtain organic matter.

Evolutionary arms race

Evolutionary arms race

In evolutionary biology, an evolutionary arms race is an ongoing struggle between competing sets of co-evolving genes, phenotypic and behavioral traits that develop escalating adaptations and counter-adaptations against each other, resembling an arms race. These are often described as examples of positive feedback. The co-evolving gene sets may be in different species, as in an evolutionary arms race between a predator species and its prey, or a parasite and its host. Alternatively, the arms race may be between members of the same species, as in the manipulation/sales resistance model of communication or as in runaway evolution or Red Queen effects. One example of an evolutionary arms race is in sexual conflict between the sexes, often described with the term Fisherian runaway. Thierry Lodé emphasized the role of such antagonistic interactions in evolution leading to character displacements and antagonistic coevolution.

Anti-predator adaptation

Anti-predator adaptation

Anti-predator adaptations are mechanisms developed through evolution that assist prey organisms in their constant struggle against predators. Throughout the animal kingdom, adaptations have evolved for every stage of this struggle, namely by avoiding detection, warding off attack, fighting back, or escaping when caught.

Consumer (food chain)

Consumer (food chain)

A consumer in a food chain is a living creature that eats organisms from a different population. A consumer is a heterotroph and a producer is an autotroph. Both are organisms that obtain energy from other living things... Like sea angels, they take in organic moles by consuming other organisms, so they are commonly called consumers. Heterotrophs can be classified by what they usually eat as herbivores, carnivores, omnivores, or decomposers. On the other hand, autotrophs are organisms that use energy directly from the sun or from chemical bonds. Autotrophs are vital to all ecosystems because all organisms need organic molecules, and only autotrophs can produce them from inorganic compounds. Autotrophs are classified as either photoautotrophs or chemoautotrophs.

Parasitoid wasp

Parasitoid wasp

Parasitoid wasps are a large group of hymenopteran superfamilies, with all but the wood wasps (Orussoidea) being in the wasp-waisted Apocrita. As parasitoids, they lay their eggs on or in the bodies of other arthropods, sooner or later causing the death of these hosts. Different species specialise in hosts from different insect orders, most often Lepidoptera, though some select beetles, flies, or bugs; the spider wasps (Pompilidae) exclusively attack spiders. More rarely, parasitoid wasps may use plant seeds as hosts, such as Torymus druparum.

Hawksbill sea turtle

Hawksbill sea turtle

The hawksbill sea turtle is a critically endangered sea turtle belonging to the family Cheloniidae. It is the only extant species in the genus Eretmochelys. The species has a global distribution, that is largely limited to tropical and subtropical marine and estuary ecosystems.

Hydrothermal vent

Hydrothermal vent

A hydrothermal vent is a fissure on the seabed from which geothermally heated water discharges. They are commonly found near volcanically active places, areas where tectonic plates are moving apart at mid-ocean ridges, ocean basins, and hotspots. Hydrothermal deposits are rocks and mineral ore deposits formed by the action of hydrothermal vents.

Diversity

Size

The blue whale is the largest animal that has ever lived.
The blue whale is the largest animal that has ever lived.

The blue whale (Balaenoptera musculus) is the largest animal that has ever lived, weighing up to 190 tonnes and measuring up to 33.6 metres (110 ft) long.[63][64][65] The largest extant terrestrial animal is the African bush elephant (Loxodonta africana), weighing up to 12.25 tonnes[63] and measuring up to 10.67 metres (35.0 ft) long.[63] The largest terrestrial animals that ever lived were titanosaur sauropod dinosaurs such as Argentinosaurus, which may have weighed as much as 73 tonnes, and Supersaurus which may have reached 39 meters.[66][67] Several animals are microscopic; some Myxozoa (obligate parasites within the Cnidaria) never grow larger than 20 µm,[68] and one of the smallest species (Myxobolus shekel) is no more than 8.5 µm when fully grown.[69]

Numbers and habitats

The following table lists estimated numbers of described extant species for all the animal groups,[70] along with their principal habitats (terrestrial, fresh water,[71] and marine),[72] and free-living or parasitic ways of life.[73] Species estimates shown here are based on numbers described scientifically; much larger estimates have been calculated based on various means of prediction, and these can vary wildly. For instance, around 25,000–27,000 species of nematodes have been described, while published estimates of the total number of nematode species include 10,000–20,000; 500,000; 10 million; and 100 million.[74] Using patterns within the taxonomic hierarchy, the total number of animal species—including those not yet described—was calculated to be about 7.77 million in 2011.[75][76][b]

Phylum Example Described species Land Sea Freshwater Free-living Parasitic
Arthropoda wasp 1,257,000[70] 1,000,000
(insects)[78]
>40,000
(Malac-
ostraca)[79]
94,000[71] Yes[72] >45,000[c][73]
Mollusca snail 85,000[70]
107,000[80]
35,000[80] 60,000[80] 5,000[71]
12,000[80]
Yes[72] >5,600[73]
Chordata green spotted frog facing right >70,000[70][81] 23,000[82] 13,000[82] 18,000[71]
9,000[82]
Yes 40
(catfish)[83][73]
Platyhelminthes Pseudoceros dimidiatus.jpg 29,500[70] Yes[84] Yes[72] 1,300[71] Yes[72]

3,000–6,500[85]

>40,000[73]

4,000–25,000[85]

Nematoda CelegansGoldsteinLabUNC.jpg 25,000[70] Yes (soil)[72] 4,000[74] 2,000[71] 11,000[74] 14,000[74]
Annelida Nerr0328.jpg 17,000[70] Yes (soil)[72] Yes[72] 1,750[71] Yes 400[73]
Cnidaria Table coral 16,000[70] Yes[72] Yes (few)[72] Yes[72] >1,350
(Myxozoa)[73]
Porifera A colourful Sponge on the Fathom.jpg 10,800[70] Yes[72] 200–300[71] Yes Yes[86]
Echinodermata Starfish, Caswell Bay - geograph.org.uk - 409413.jpg 7,500[70] 7,500[70] Yes[72]
Bryozoa Bryozoan at Ponta do Ouro, Mozambique (6654415783).jpg 6,000[70] Yes[72] 60–80[71] Yes
Rotifera 20090730 020239 Rotifer.jpg 2,000[70] >400[87] 2,000[71] Yes
Nemertea Némerte.jpg 1,350[88][89] Yes Yes Yes
Tardigrada Tardigrade (50594282802).jpg 1,335[70] Yes[90]
(moist plants)
Yes Yes Yes
Gastrotricha Paradasys subterraneus.jpg 794[70] Yes[90] Yes Yes
Xenacoelomorpha Proporus sp.png 430[70] Yes[90] Yes
Nematomorpha Nematomorpha Somiedo.JPG 354[70] Yes
(moist places)[90]
Yes
(one genus)[91]
Yes Yes
(as adults)[90]
Yes
(as juveniles)[90]
Brachiopoda Liospiriferina rostrata Noir.jpg 396[70]
(30,000 extinct)[90]
Yes[90] Yes
Kinorhyncha Cephalorhyncha flosculosa zoomed.jpg 196[70] Yes (mud)[90] Yes
Ctenophora Bathocyroe fosteri.jpg 187[70] Yes[90] Yes
Onychophora Unidentified velvet worm.jpg 187[70] Yes[90] Yes
Chaetognatha Chaetoblack.png 186[70] Yes[90] Yes
Entoprocta Barentsia laxa 1498966.png 172[70] Yes[90] Yes
Yes
Hemichordata Torq ventral acorn worm.tif 126[70] Yes[90] Yes
Rhombozoa Dicyema japonicum.png 107[70] Yes
Gnathostomulida Gnathostomula paradoxa Sylt.tif 97[70] Yes (sand)[90] Yes
Loricifera Pliciloricus enigmatus.jpg 30[70] Yes (sand)[90] Yes
Orthonectida EB1911 Mesozoa - Rhopalura giardii.jpg 29[70] Yes
Priapulida Halicryptus spinulosus 1.JPEG 20[70] Yes[90] Yes
Phoronida Phoronis ijimai 99523588.jpg 16[70] Yes[90] Yes
Placozoa Trichoplax adhaerens photograph.png 4[92] Yes[90] Yes
Cycliophora Feeding stage and attached Prometheus larva of Symbion pandora.jpg 2[93] Yes[93] Yes[94][93]
Micrognathozoa Limnognathia maerski youtube.png One[90] Yes (sand)[90] Yes
Total number of described extant species as of 2013: 1,525,728[70]

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Largest organisms

Largest organisms

The largest organisms now found on Earth can be determined according to various aspects of an organism's size, such as: mass, volume, area, length, height, or even genome size. Some organisms group together to form a superorganism, but such are not classed as single large organisms. The Great Barrier Reef is the world's largest structure composed of living entities, stretching 2,000 km (1,200 mi), but contains many organisms of many types of species.

Smallest organisms

Smallest organisms

The smallest organisms found on Earth can be determined according to various aspects of organism size, including volume, mass, height, length, or genome size.

Blue whale

Blue whale

The blue whale is a marine mammal and a baleen whale. Reaching a maximum confirmed length of 29.9 meters (98 ft) and weighing up to 199 tonnes, it is the largest animal known ever to have existed. The blue whale's long and slender body can be of various shades of greyish-blue dorsally and somewhat lighter underneath. Four subspecies are recognized: B. m. musculus in the North Atlantic and North Pacific, B. m. intermedia in the Southern Ocean, B. m. brevicauda in the Indian Ocean and South Pacific Ocean, B. m. indica in the Northern Indian Ocean. There is also a population in the waters off Chile that may constitute a fifth subspecies.

African bush elephant

African bush elephant

The African bush elephant, also known as the African savanna elephant, is one of two extant African elephant species and one of three extant elephant species. It is the largest living terrestrial animal, with bulls reaching a shoulder height of up to 3.96 m and a body mass of up to 10.4 t.

Sauropoda

Sauropoda

Sauropoda, whose members are known as sauropods, is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads, and four thick, pillar-like legs. They are notable for the enormous sizes attained by some species, and the group includes the largest animals to have ever lived on land. Well-known genera include Brachiosaurus, Diplodocus, Apatosaurus and Brontosaurus.

Argentinosaurus

Argentinosaurus

Argentinosaurus is a genus of giant sauropod dinosaur that lived during the Late Cretaceous period in what is now Argentina. Although it is only known from fragmentary remains, Argentinosaurus is one of the largest known land animals of all time, perhaps the largest, reaching 30–35 metres (98–115 ft) in length and 60–75 tonnes in body mass. It was a member of Titanosauria, the dominant group of sauropods during the Cretaceous. It is widely regarded by many paleontologists as the biggest dinosaur ever, and perhaps lengthwise the longest animal ever, though both claims have no concrete evidence yet.

Myxozoa

Myxozoa

Myxozoa is a subphylum of aquatic cnidarian animals – all obligate parasites. It contains the smallest animals ever known to have lived. Over 2,180 species have been described and some estimates have suggested at least 30,000 undiscovered species. Many have a two-host lifecycle, involving a fish and an annelid worm or a bryozoan. The average size of a myxosporean spore usually ranges from 10 μm to 20 μm, whereas that of a malacosporean spore can be up to 2 mm. Myxozoans can live in both freshwater and marine habitats.

Obligate parasite

Obligate parasite

An obligate parasite or holoparasite is a parasitic organism that cannot complete its life-cycle without exploiting a suitable host. If an obligate parasite cannot obtain a host it will fail to reproduce. This is opposed to a facultative parasite, which can act as a parasite but does not rely on its host to continue its life-cycle. Obligate parasites have evolved a variety of parasitic strategies to exploit their hosts. Holoparasites and some hemiparasites are obligate.

Micrometre

Micrometre

The micrometre or micrometer, also commonly known as a micron, is a unit of length in the International System of Units (SI) equalling 1×10−6 metre ; that is, one millionth of a metre.

Taxonomy (biology)

Taxonomy (biology)

In biology, taxonomy is the scientific study of naming, defining (circumscribing) and classifying groups of biological organisms based on shared characteristics. Organisms are grouped into taxa and these groups are given a taxonomic rank; groups of a given rank can be aggregated to form a more inclusive group of higher rank, thus creating a taxonomic hierarchy. The principal ranks in modern use are domain, kingdom, phylum, class, order, family, genus, and species. The Swedish botanist Carl Linnaeus is regarded as the founder of the current system of taxonomy, as he developed a ranked system known as Linnaean taxonomy for categorizing organisms and binomial nomenclature for naming organisms.

Phylum

Phylum

In biology, a phylum is a level of classification or taxonomic rank below kingdom and above class. Traditionally, in botany the term division has been used instead of phylum, although the International Code of Nomenclature for algae, fungi, and plants accepts the terms as equivalent. Depending on definitions, the animal kingdom Animalia contains about 31 phyla, the plant kingdom Plantae contains about 14 phyla, and the fungus kingdom Fungi contains about 8 phyla. Current research in phylogenetics is uncovering the relationships between phyla, which are contained in larger clades, like Ecdysozoa and Embryophyta.

Mollusca

Mollusca

Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks. Around 85,000 extant species of molluscs are recognized. The number of fossil species is estimated between 60,000 and 100,000 additional species. The proportion of undescribed species is very high. Many taxa remain poorly studied.

Evolutionary origin

Animals are found as long ago as the Ediacaran biota, towards the end of the Precambrian, and possibly somewhat earlier. It had long been doubted whether these life-forms included animals,[95][96][97] but the discovery of the animal lipid cholesterol in fossils of Dickinsonia establishes their nature.[98] Animals are thought to have originated under low-oxygen conditions, suggesting that they were capable of living entirely by anaerobic respiration, but as they became specialized for aerobic metabolism they became fully dependent on oxygen in their environments.[99]

Many animal phyla first appear in the fossil record during the Cambrian explosion, starting about 539 million years ago, in beds such as the Burgess shale.[100] Extant phyla in these rocks include molluscs, brachiopods, onychophorans, tardigrades, arthropods, echinoderms and hemichordates, along with numerous now-extinct forms such as the predatory Anomalocaris. The apparent suddenness of the event may however be an artefact of the fossil record, rather than showing that all these animals appeared simultaneously.[101][102][103][104] That view is supported by the discovery of Auroralumina attenboroughii, the earliest known Ediacaran crown-group cnidarian (557–562 mya, some 20 million years before the Cambrian explosion) from Charnwood Forest, England. It is thought to be one of the earliest predators, catching small prey with its nematocysts as modern cnidarians do.[105]

Some palaeontologists have suggested that animals appeared much earlier than the Cambrian explosion, possibly as early as 1 billion years ago.[106] Early fossils that might represent animals appear for example in the 665-million-year-old rocks of the Trezona Formation of South Australia. These fossils are interpreted as most probably being early sponges.[107] Trace fossils such as tracks and burrows found in the Tonian period (from 1 gya) may indicate the presence of triploblastic worm-like animals, roughly as large (about 5 mm wide) and complex as earthworms.[108] However, similar tracks are produced today by the giant single-celled protist Gromia sphaerica, so the Tonian trace fossils may not indicate early animal evolution.[109][110] Around the same time, the layered mats of microorganisms called stromatolites decreased in diversity, perhaps due to grazing by newly evolved animals.[111] Objects such as sediment-filled tubes that resemble trace fossils of the burrows of wormlike animals have been found in 1.2 gya rocks in North America, in 1.5 gya rocks in Australia and North America, and in 1.7 gya rocks in Australia. Their interpretation as having an animal origin is disputed, as they might be water-escape or other structures.[112][113]

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Ediacaran biota

Ediacaran biota

The Ediacaran biota is a taxonomic period classification that consists of all life forms that were present on Earth during the Ediacaran Period. These were composed of enigmatic tubular and frond-shaped, mostly sessile, organisms. Trace fossils of these organisms have been found worldwide, and represent the earliest known complex multicellular organisms. The term "Ediacara biota" has received criticism from some scientists due to its alleged inconsistency, arbitrary exclusion of certain fossils, and inability to be precisely defined.

Cholesterol

Cholesterol

Cholesterol is any of a class of certain organic molecules called lipids. It is a sterol, a type of lipid. Cholesterol is biosynthesized by all animal cells and is an essential structural component of animal cell membranes. When chemically isolated, it is a yellowish crystalline solid.

Dickinsonia

Dickinsonia

Dickinsonia is an extinct genus of basal animal that lived during the late Ediacaran period in what is now Australia, China, Russia and Ukraine. The individual Dickinsonia typically resembles a bilaterally symmetrical ribbed oval. Its affinities are presently unknown; its mode of growth is consistent with a stem-group bilaterian affinity, though some have suggested that it belongs to the fungi, or even an "extinct kingdom". The discovery of cholesterol molecules in fossils of Dickinsonia lends support to the idea that Dickinsonia was an animal.

Anaerobic respiration

Anaerobic respiration

Anaerobic respiration is respiration using electron acceptors other than molecular oxygen (O2). Although oxygen is not the final electron acceptor, the process still uses a respiratory electron transport chain.

Fossil

Fossil

A fossil is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved in amber, hair, petrified wood and DNA remnants. The totality of fossils is known as the fossil record.

Cambrian explosion

Cambrian explosion

The Cambrian explosion, Cambrian radiation, Cambrian diversification, or the Biological Big Bang refers to an interval of time approximately 538.8 million years ago in the Cambrian Period when practically all major animal phyla started appearing in the fossil record. It lasted for about 13 – 25 million years and resulted in the divergence of most modern metazoan phyla. The event was accompanied by major diversification in other groups of organisms as well.

Brachiopod

Brachiopod

Brachiopods, phylum Brachiopoda, are a phylum of trochozoan animals that have hard "valves" (shells) on the upper and lower surfaces, unlike the left and right arrangement in bivalve molluscs. Brachiopod valves are hinged at the rear end, while the front can be opened for feeding or closed for protection. Two major categories are traditionally recognized, articulate and inarticulate brachiopods. The word "articulate" is used to describe the tooth-and-groove structures of the valve-hinge which is present in the articulate group, and absent from the inarticulate group. This is the leading diagnostic skeletal feature, by which the two main groups can be readily distinguished as fossils. Articulate brachiopods have toothed hinges and simple, vertically-oriented opening and closing muscles. Conversely, inarticulate brachiopods have weak, untoothed hinges and a more complex system of vertical and oblique (diagonal) muscles used to keep the two valves aligned. In many brachiopods, a stalk-like pedicle projects from an opening near the hinge of one of the valves, known as the pedicle or ventral valve. The pedicle, when present, keeps the animal anchored to the seabed but clear of sediment which would obstruct the opening.

Arthropod

Arthropod

Arthropods are invertebrate animals with an exoskeleton, a segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and cuticle made of chitin, often mineralised with calcium carbonate. The arthropod body plan consists of segments, each with a pair of appendages. Arthropods are bilaterally symmetrical and their body possesses an external skeleton. In order to keep growing, they must go through stages of moulting, a process by which they shed their exoskeleton to reveal a new one. Some species have wings. They are an extremely diverse group, with up to 10 million species.

Echinoderm

Echinoderm

An echinoderm is any member of the phylum Echinodermata. The adults are recognisable by their radial symmetry, and include starfish, brittle stars, sea urchins, sand dollars, and sea cucumbers, as well as the sea lilies or "stone lilies". Adult echinoderms are found on the sea bed at every ocean depth, from the intertidal zone to the abyssal zone. The phylum contains about 7,000 living species, making it the second-largest grouping of deuterostomes, after the chordates. Echinoderms are the largest entirely marine phylum. The first definitive echinoderms appeared near the start of the Cambrian.

Hemichordate

Hemichordate

Hemichordata is a phylum which consists of triploblastic, enterocoelomate, and bilaterally symmetrical marine deuterostome animals, generally considered the sister group of the echinoderms. They appear in the Lower or Middle Cambrian and include two main classes: Enteropneusta, and Pterobranchia. A third class, Planctosphaeroidea, is known only from the larva of a single species, Planctosphaera pelagica. The class Graptolithina, formerly considered extinct, is now placed within the pterobranchs, represented by a single living genus Rhabdopleura.

Anomalocaris

Anomalocaris

Anomalocaris is an extinct genus of radiodont, an order of early-diverging stem-group arthropods. The first fossils of Anomalocaris were discovered in the Ogygopsis Shale of the Stephen Formation in British Columbia, Canada by Joseph Frederick Whiteaves, with more examples found by Charles Doolittle Walcott in the Burgess Shale unit of the Stephen Formation. Other closely related fossils have been found in the older Emu Bay Shale of Australia, as well as possibly elsewhere. Originally several fossilized parts discovered separately were thought to be three separate creatures, a misapprehension corrected by Harry B. Whittington and Derek Briggs in a 1985 journal article. With a body length close to 40 centimetres, A. canadensis is thought to be one of the earliest examples of an apex predator, though others have been found in older Cambrian lagerstätten deposits.

Charnwood Forest

Charnwood Forest

Charnwood Forest is a hilly tract in north-western Leicestershire, England, bounded by Leicester, Loughborough and Coalville. The area is undulating, rocky and picturesque, with barren areas. It also has some extensive tracts of woodland; its elevation is generally 600 feet (180 m) and upwards, the area exceeding this height being about 6,100 acres (25 km2). The highest point, Bardon Hill, is 912 feet (278 m). On its western flank lies an abandoned coalfield, with Coalville and other former mining villages, now being regenerated and replanted as part of the National Forest. The M1 motorway, between junctions 22 and 23, cuts through Charnwood Forest.

Phylogeny

Animals are monophyletic, meaning they are derived from a common ancestor. Animals are sister to the Choanoflagellata, with which they form the Choanozoa.[116] The most basal animals, the Porifera, Ctenophora, Cnidaria, and Placozoa, have body plans that lack bilateral symmetry. Their relationships are still disputed; the sister group to all other animals could be the Porifera or the Ctenophora,[117] both of which lack hox genes, important in body plan development.[118]

These genes are found in the Placozoa[119][120] and the higher animals, the Bilateria.[121][122] 6,331 groups of genes common to all living animals have been identified; these may have arisen from a single common ancestor that lived 650 million years ago in the Precambrian. 25 of these are novel core gene groups, found only in animals; of those, 8 are for essential components of the Wnt and TGF-beta signalling pathways which may have enabled animals to become multicellular by providing a pattern for the body's system of axes (in three dimensions), and another 7 are for transcription factors including homeodomain proteins involved in the control of development.[123][124]

The phylogenetic tree indicates approximately how many millions of years ago (mya) the lineages split.[125][126][127][128][129]

Choanozoa

Choanoflagellata Desmarella moniliformis.jpg

Animalia
Parazoa

Porifera Reef3859 - Flickr - NOAA Photo Library.jpg

Eumetazoa

Ctenophora Comb jelly.jpg

ParaHoxozoa

Placozoa Trichoplax adhaerens photograph.png

Cnidaria Cauliflour Jellyfish, Cephea cephea at Marsa Shouna, Red Sea, Egypt SCUBA.jpg

Bilateria

Xenacoelomorpha Proporus sp.png

Nephrozoa
Deuterostomia

Chordata Common carp (white background).jpg

Ambulacraria Portugal 20140812-DSC01434 (21371237591).jpg

Protostomia
Ecdysozoa

Scalidophora Priapulus caudatus 20150625.jpg

Panarthropoda Long nosed weevil edit.jpg

Nematoida CelegansGoldsteinLabUNC 2.jpg

>529 mya
Spiralia
Gnathifera

Chaetognatha Chaetoblack 3.png

Gnathostomulida

Micrognathozoa

Rotifera (inc. Acanthocephala) Bdelloid Rotifer (cropped).jpg

Platytrochozoa
Rouphozoa

Gastrotricha

Platyhelminthes Sorocelis reticulosa.jpg

Lophotrochozoa

Annelida Polychaeta (no) 2.jpg

Mollusca Grapevinesnail 01.jpg

Nemertea Amphiporus angulatus.png

Lophophorata

Bryozoa

Brachiozoa

Brachiopoda

Phoronida

550 mya
580 mya
610 mya
650 mya
Triploblasts
680 mya
760 mya
950 mya

Non-bilateria

Non-bilaterians include sponges (centre) and corals (background).
Non-bilaterians include sponges (centre) and corals (background).

Several animal phyla lack bilateral symmetry. Among these, the sponges (Porifera) probably diverged first, representing the oldest animal phylum.[130] Sponges lack the complex organization found in most other animal phyla;[131] their cells are differentiated, but in most cases not organised into distinct tissues.[132] They typically feed by drawing in water through pores.[133]

The Ctenophora (comb jellies) and Cnidaria (which includes jellyfish, sea anemones, and corals) are radially symmetric and have digestive chambers with a single opening, which serves as both mouth and anus.[134] They are sometimes placed together in the group Coelenterata because of common traits, not because of close relationships. Animals in both phyla have distinct tissues, but these are not organised into organs.[135] They are diploblastic, having only two main germ layers, ectoderm and endoderm.[136] The tiny placozoans are similar, but they do not have a permanent digestive chamber.[137][138]

Bilateria

Idealised bilaterian body plan.[d] With an elongated body and a direction of movement the animal has head and tail ends. Sense organs and mouth form the basis of the head. Opposed circular and longitudinal muscles enable peristaltic motion.
Idealised bilaterian body plan.[d] With an elongated body and a direction of movement the animal has head and tail ends. Sense organs and mouth form the basis of the head. Opposed circular and longitudinal muscles enable peristaltic motion.

The remaining animals, the great majority—comprising some 29 phyla and over a million species—form a clade, the Bilateria, which have a bilaterally symmetric body plan. The Bilateria are triploblastic, with three well-developed germ layers, and their tissues form distinct organs. The digestive chamber has two openings, a mouth and an anus, and there is an internal body cavity, a coelom or pseudocoelom. These animals have a head end (anterior) and a tail end (posterior), a back (dorsal) surface and a belly (ventral) surface, and a left and a right side.[139][140]

Having a front end means that this part of the body encounters stimuli, such as food, favouring cephalisation, the development of a head with sense organs and a mouth. Many bilaterians have a combination of circular muscles that constrict the body, making it longer, and an opposing set of longitudinal muscles, that shorten the body;[140] these enable soft-bodied animals with a hydrostatic skeleton to move by peristalsis.[141] They also have a gut that extends through the basically cylindrical body from mouth to anus. Many bilaterian phyla have primary larvae which swim with cilia and have an apical organ containing sensory cells. However, over evolutionary time, descendant spaces have evolved which have lost one or more of each of these characteristics. For example, adult echinoderms are radially symmetric (unlike their larvae), while some parasitic worms have extremely simplified body structures.[139][140]

Genetic studies have considerably changed zoologists' understanding of the relationships within the Bilateria. Most appear to belong to two major lineages, the protostomes and the deuterostomes.[142] The basalmost bilaterians are the Xenacoelomorpha.[143][144][145]

Protostomes and deuterostomes

The bilaterian gut develops in two ways. In many protostomes, the blastopore develops into the mouth, while in deuterostomes it becomes the anus.
The bilaterian gut develops in two ways. In many protostomes, the blastopore develops into the mouth, while in deuterostomes it becomes the anus.

Protostomes and deuterostomes differ in several ways. Early in development, deuterostome embryos undergo radial cleavage during cell division, while many protostomes (the Spiralia) undergo spiral cleavage.[146] Animals from both groups possess a complete digestive tract, but in protostomes the first opening of the embryonic gut develops into the mouth, and the anus forms secondarily. In deuterostomes, the anus forms first while the mouth develops secondarily.[147][148] Most protostomes have schizocoelous development, where cells simply fill in the interior of the gastrula to form the mesoderm. In deuterostomes, the mesoderm forms by enterocoelic pouching, through invagination of the endoderm.[149]

The main deuterostome phyla are the Echinodermata and the Chordata.[150] Echinoderms are exclusively marine and include starfish, sea urchins, and sea cucumbers.[151] The chordates are dominated by the vertebrates (animals with backbones),[152] which consist of fishes, amphibians, reptiles, birds, and mammals.[153] The deuterostomes also include the Hemichordata (acorn worms).[154][155]

Ecdysozoa
Ecdysis: a dragonfly has emerged from its dry exuviae and is expanding its wings. Like other arthropods, its body is divided into segments.
Ecdysis: a dragonfly has emerged from its dry exuviae and is expanding its wings. Like other arthropods, its body is divided into segments.

The Ecdysozoa are protostomes, named after their shared trait of ecdysis, growth by moulting.[156] They include the largest animal phylum, the Arthropoda, which contains insects, spiders, crabs, and their kin. All of these have a body divided into repeating segments, typically with paired appendages. Two smaller phyla, the Onychophora and Tardigrada, are close relatives of the arthropods and share these traits. The ecdysozoans also include the Nematoda or roundworms, perhaps the second largest animal phylum. Roundworms are typically microscopic, and occur in nearly every environment where there is water;[157] some are important parasites.[158] Smaller phyla related to them are the Nematomorpha or horsehair worms, and the Kinorhyncha, Priapulida, and Loricifera. These groups have a reduced coelom, called a pseudocoelom.[159]

Spiralia
Spiral cleavage in a sea snail embryo
Spiral cleavage in a sea snail embryo

The Spiralia are a large group of protostomes that develop by spiral cleavage in the early embryo.[160] The Spiralia's phylogeny has been disputed, but it contains a large clade, the superphylum Lophotrochozoa, and smaller groups of phyla such as the Rouphozoa which includes the gastrotrichs and the flatworms. All of these are grouped as the Platytrochozoa, which has a sister group, the Gnathifera, which includes the rotifers.[161][162]

The Lophotrochozoa includes the molluscs, annelids, brachiopods, nemerteans, bryozoa and entoprocts.[161][163][164] The molluscs, the second-largest animal phylum by number of described species, includes snails, clams, and squids, while the annelids are the segmented worms, such as earthworms, lugworms, and leeches. These two groups have long been considered close relatives because they share trochophore larvae.[165][166]

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Lists of animals

Lists of animals

Animals are a form of eukaryotes in the biological kingdom Animalia. With few exceptions, animals consume organic material, breathe oxygen, are able to move, reproduce sexually, and grow from a hollow sphere of cells, the blastula, during embryonic development. Over 1.5 million living animal species have been described—of which around 1 million are insects—but it has been estimated there are over 7 million in total. Animals range in size from 8.5 millionths of a metre to 33.6 metres (110 ft) long and have complex interactions with each other and their environments, forming intricate food webs. The study of animals is called zoology.

Monophyly

Monophyly

In biological cladistics for the classification of organisms, monophyly is the condition of a taxonomic grouping being a clade — that is, a grouping of taxa which meets these criteria:the grouping contains its own most recent common ancestor, i.e. excludes non-descendants of that common ancestor the grouping contains all the descendants of that common ancestor, without exception

Choanozoa

Choanozoa

Choanozoa is a clade of opisthokont eukaryotes consisting of the choanoflagellates (Choanoflagellatea) and the animals. The sister-group relationship between the choanoflagellates and animals has important implications for the origin of the animals. The clade was identified in 2015 by Graham Budd and Sören Jensen, who used the name Apoikozoa. The 2018 revision of the classification first proposed by the International Society of Protistologists in 2012 recommends the use of the name Choanozoa.

Ctenophora

Ctenophora

Ctenophora comprise a phylum of marine invertebrates, commonly known as comb jellies, that inhabit sea waters worldwide. They are notable for the groups of cilia they use for swimming, and they are the largest animals to swim with the help of cilia.

Cnidaria

Cnidaria

Cnidaria is a phylum under kingdom Animalia containing over 11,000 species of aquatic animals found both in freshwater and marine environments, predominantly the latter.

Placozoa

Placozoa

Placozoa is a phylum that consists of a diverse group of basal form of marine free-living (non-parasitic) multicellular organism called placozoans. They are the simplest in structure of all animals. Three genera have been found: the classical Trichoplax adhaerens, Hoilungia hongkongensis, and Polyplacotoma mediterranea, where the last appears most basal. The last two have been found only since 2017. Although the Placozoa were first discovered in 1883 by the German zoologist Franz Eilhard Schulze (1840–1921) and since the 1970s more systematically analyzed by the German protozoologist Karl Gottlieb Grell (1912–1994), a common name does not yet exist for the taxon; the scientific name means "flat animals".

Symmetry in biology

Symmetry in biology

Symmetry in biology refers to the symmetry observed in organisms, including plants, animals, fungi, and bacteria. External symmetry can be easily seen by just looking at an organism. For example, take the face of a human being which has a plane of symmetry down its centre, or a pine cone with a clear symmetrical spiral pattern. Internal features can also show symmetry, for example the tubes in the human body which are cylindrical and have several planes of symmetry.

Hox gene

Hox gene

Hox genes, a subset of homeobox genes, are a group of related genes that specify regions of the body plan of an embryo along the head-tail axis of animals. Hox proteins encode and specify the characteristics of 'position', ensuring that the correct structures form in the correct places of the body. For example, Hox genes in insects specify which appendages form on a segment, and Hox genes in vertebrates specify the types and shape of vertebrae that will form. In segmented animals, Hox proteins thus confer segmental or positional identity, but do not form the actual segments themselves.

Gene

Gene

In biology, the word gene can have several different meanings. The Mendelian gene is a basic unit of heredity and the molecular gene is a sequence of nucleotides in DNA that is transcribed to produce a functional RNA. There are two types of molecular genes: protein-coding genes and noncoding genes.

Cryogenian

Cryogenian

The Cryogenian is a geologic period that lasted from 720 to 635 million years ago. It forms the second geologic period of the Neoproterozoic Era, preceded by the Tonian Period and followed by the Ediacaran.

Precambrian

Precambrian

The Precambrian is the earliest part of Earth's history, set before the current Phanerozoic Eon. The Precambrian is so named because it preceded the Cambrian, the first period of the Phanerozoic Eon, which is named after Cambria, the Latinised name for Wales, where rocks from this age were first studied. The Precambrian accounts for 88% of the Earth's geologic time.

Evo-devo gene toolkit

Evo-devo gene toolkit

The evo-devo gene toolkit is the small subset of genes in an organism's genome whose products control the organism's embryonic development. Toolkit genes are central to the synthesis of molecular genetics, palaeontology, evolution and developmental biology in the science of evolutionary developmental biology (evo-devo). Many of them are ancient and highly conserved among animal phyla.

History of classification

Jean-Baptiste de Lamarck led the creation of a modern classification of invertebrates, breaking up Linnaeus's "Vermes" into 9 phyla by 1809.[167]
Jean-Baptiste de Lamarck led the creation of a modern classification of invertebrates, breaking up Linnaeus's "Vermes" into 9 phyla by 1809.[167]

In the classical era, Aristotle divided animals,[e] based on his own observations, into those with blood (roughly, the vertebrates) and those without. The animals were then arranged on a scale from man (with blood, 2 legs, rational soul) down through the live-bearing tetrapods (with blood, 4 legs, sensitive soul) and other groups such as crustaceans (no blood, many legs, sensitive soul) down to spontaneously generating creatures like sponges (no blood, no legs, vegetable soul). Aristotle was uncertain whether sponges were animals, which in his system ought to have sensation, appetite, and locomotion, or plants, which did not: he knew that sponges could sense touch, and would contract if about to be pulled off their rocks, but that they were rooted like plants and never moved about.[168]

In 1758, Carl Linnaeus created the first hierarchical classification in his Systema Naturae.[169] In his original scheme, the animals were one of three kingdoms, divided into the classes of Vermes, Insecta, Pisces, Amphibia, Aves, and Mammalia. Since then the last four have all been subsumed into a single phylum, the Chordata, while his Insecta (which included the crustaceans and arachnids) and Vermes have been renamed or broken up. The process was begun in 1793 by Jean-Baptiste de Lamarck, who called the Vermes une espèce de chaos (a chaotic mess)[f] and split the group into three new phyla: worms, echinoderms, and polyps (which contained corals and jellyfish). By 1809, in his Philosophie Zoologique, Lamarck had created 9 phyla apart from vertebrates (where he still had 4 phyla: mammals, birds, reptiles, and fish) and molluscs, namely cirripedes, annelids, crustaceans, arachnids, insects, worms, radiates, polyps, and infusorians.[167]

In his 1817 Le Règne Animal, Georges Cuvier used comparative anatomy to group the animals into four embranchements ("branches" with different body plans, roughly corresponding to phyla), namely vertebrates, molluscs, articulated animals (arthropods and annelids), and zoophytes (radiata) (echinoderms, cnidaria and other forms).[171] This division into four was followed by the embryologist Karl Ernst von Baer in 1828, the zoologist Louis Agassiz in 1857, and the comparative anatomist Richard Owen in 1860.[172]

In 1874, Ernst Haeckel divided the animal kingdom into two subkingdoms: Metazoa (multicellular animals, with five phyla: coelenterates, echinoderms, articulates, molluscs, and vertebrates) and Protozoa (single-celled animals), including a sixth animal phylum, sponges.[173][172] The protozoa were later moved to the former kingdom Protista, leaving only the Metazoa as a synonym of Animalia.[174]

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Taxonomy (biology)

Taxonomy (biology)

In biology, taxonomy is the scientific study of naming, defining (circumscribing) and classifying groups of biological organisms based on shared characteristics. Organisms are grouped into taxa and these groups are given a taxonomic rank; groups of a given rank can be aggregated to form a more inclusive group of higher rank, thus creating a taxonomic hierarchy. The principal ranks in modern use are domain, kingdom, phylum, class, order, family, genus, and species. The Swedish botanist Carl Linnaeus is regarded as the founder of the current system of taxonomy, as he developed a ranked system known as Linnaean taxonomy for categorizing organisms and binomial nomenclature for naming organisms.

Aristotle's biology

Aristotle's biology

Aristotle's biology is the theory of biology, grounded in systematic observation and collection of data, mainly zoological, embodied in Aristotle's books on the science. Many of his observations were made during his stay on the island of Lesbos, including especially his descriptions of the marine biology of the Pyrrha lagoon, now the Gulf of Kalloni. His theory is based on his concept of form, which derives from but is markedly unlike Plato's theory of Forms.

Aristotle

Aristotle

Aristotle was an Ancient Greek philosopher and polymath. His writings cover a broad range of subjects including physics, biology, zoology, metaphysics, logic, ethics, aesthetics, poetry, drama, music, rhetoric, psychology, linguistics, economics, politics, meteorology, geology, and government. As the founder of the Peripatetic school of philosophy in the Lyceum in Athens, he began the wider Aristotelian tradition that followed, which set the groundwork for the development of modern science.

Carl Linnaeus

Carl Linnaeus

Carl Linnaeus, also known after his ennoblement in 1761 as Carl von Linné, was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the modern system of naming organisms. He is known as the "father of modern taxonomy". Many of his writings were in Latin; his name is rendered in Latin as Carolus Linnæus and, after his 1761 ennoblement, as Carolus a Linné.

Systema Naturae

Systema Naturae

Systema Naturae is one of the major works of the Swedish botanist, zoologist and physician Carl Linnaeus (1707–1778) and introduced the Linnaean taxonomy. Although the system, now known as binomial nomenclature, was partially developed by the Bauhin brothers, Gaspard and Johann, Linnaeus was first to use it consistently throughout his book. The first edition was published in 1735. The full title of the 10th edition (1758), which was the most important one, was Systema naturæ per regna tria naturæ, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis or translated: "System of nature through the three kingdoms of nature, according to classes, orders, genera and species, with characters, differences, synonyms, places".

Vermes in the 10th edition of Systema Naturae

Vermes in the 10th edition of Systema Naturae

In 1758, in the 10th edition of Systema Naturae, the Swedish scientist and taxonomist Carl Linnaeus described the class "Vermes" as:Animals of slow motion, soft substance, able to increase their bulk and restore parts which have been destroyed, extremely tenacious of life, and the inhabitants of moist places. Many of them are without a distinct head, and most of them without feet. They are principally distinguished by their tentacles. By the Ancients they were not improperly called imperfect animals, as being destitute of ears, nose, head, eyes and legs; and are therefore totally distinct from Insects.

Insecta in the 10th edition of Systema Naturae

Insecta in the 10th edition of Systema Naturae

In the 10th edition of Systema Naturae, Carl Linnaeus classified the arthropods, including insects, arachnids and crustaceans, among his class "Insecta". He described the Insecta as:A very numerous and various class consisting of small animals, breathing through lateral spiracles, armed on all sides with a bony skin, or covered with hair; furnished with many feet, and moveable antennae, which project from the head, and are the probable instruments of sensation.

Pisces in the 10th edition of Systema Naturae

Pisces in the 10th edition of Systema Naturae

In the 10th edition of Systema Naturae, Carl Linnaeus described the Pisces as:Always inhabiting the waters; are swift in their motion and voracious in their appetites. They breathe by means of gills, which are generally united by a bony arch; swim by means of radiate fins, and are mostly covered over with cartilaginous scales. Besides the parts they have in common with other animals, they are furnished with a nictitant membrane, and most of them with a swim-bladder, by the contraction or dilatation of which, they can raise or sink themselves in their element at pleasure.

Amphibia in the 10th edition of Systema Naturae

Amphibia in the 10th edition of Systema Naturae

In the 10th edition of Systema Naturae, Carl Linnaeus described the Amphibia as:Animals that are distinguished by a body cold and generally naked; stern and expressive countenance; harsh voice; mostly lurid color; filthy odor; a few are furnished with a horrid poison; all have cartilaginous bones, slow circulation, exquisite sight and hearing, large pulmonary vessels, lobate liver, oblong thick stomach, and cystic, hepatic, and pancreatic ducts: they are deficient in diaphragm, do not transpire (sweat), can live a long time without food, are tenatious of life, and have the power of reproducing parts which have been destroyed or lost; some undergo a metamorphosis; some cast (shed) their skin; some appear to live promiscuously on land or in the water, and some are torpid during the winter.

Aves in the 10th edition of Systema Naturae

Aves in the 10th edition of Systema Naturae

In the 10th edition of Systema Naturae, published in 1758, the Swedish naturalist Carl Linnaeus described 554 species of bird and gave each a binomial name.

Mammalia in the 10th edition of Systema Naturae

Mammalia in the 10th edition of Systema Naturae

The Mammalia in the 10th edition of Systema Naturae forms one of six classes of animals in Carl Linnaeus's tenth reformed edition written in Latin. The following explanations are based on William Turton's translations who rearranged and corrected earlier editions published by Johann Friedrich Gmelin, Johan Christian Fabricius and Carl Ludwig Willdenow:Animals that suckle their young by means of lactiferous teats. In external and internal structure they resemble man: most of them are quadrupeds; and with man, their natural enemy, inhabit the surface of the Earth. The largest, though fewest in number, inhabit the ocean.

Chordate

Chordate

A chordate is an animal of the phylum Chordata. All chordates possess, at some point during their larval or adult stages, five synapomorphies, or primary physical characteristics, that distinguish them from all the other taxa. These five synapomorphies include a notochord, dorsal hollow nerve cord, endostyle or thyroid, pharyngeal slits, and a post-anal tail. The name “chordate” comes from the first of these synapomorphies, the notochord, which plays a significant role in chordate structure and movement. Chordates are also bilaterally symmetric, have a coelom, possess a circulatory system, and exhibit metameric segmentation.

In human culture

Practical uses

Sides of beef in a slaughterhouse
Sides of beef in a slaughterhouse

The human population exploits a large number of other animal species for food, both of domesticated livestock species in animal husbandry and, mainly at sea, by hunting wild species.[175][176] Marine fish of many species are caught commercially for food. A smaller number of species are farmed commercially.[175][177][178] Humans and their livestock make up more than 90% of the biomass of all terrestrial vertebrates, and almost as much as all insects combined.[179]

Invertebrates including cephalopods, crustaceans, and bivalve or gastropod molluscs are hunted or farmed for food.[180] Chickens, cattle, sheep, pigs, and other animals are raised as livestock for meat across the world.[176][181][182] Animal fibres such as wool are used to make textiles, while animal sinews have been used as lashings and bindings, and leather is widely used to make shoes and other items. Animals have been hunted and farmed for their fur to make items such as coats and hats.[183] Dyestuffs including carmine (cochineal),[184][185] shellac,[186][187] and kermes[188][189] have been made from the bodies of insects. Working animals including cattle and horses have been used for work and transport from the first days of agriculture.[190]

Animals such as the fruit fly Drosophila melanogaster serve a major role in science as experimental models.[191][192][193][194] Animals have been used to create vaccines since their discovery in the 18th century.[195] Some medicines such as the cancer drug Yondelis are based on toxins or other molecules of animal origin.[196]

A gun dog retrieving a duck during a hunt
A gun dog retrieving a duck during a hunt

People have used hunting dogs to help chase down and retrieve animals,[197] and birds of prey to catch birds and mammals,[198] while tethered cormorants have been used to catch fish.[199] Poison dart frogs have been used to poison the tips of blowpipe darts.[200][201] A wide variety of animals are kept as pets, from invertebrates such as tarantulas and octopuses, insects including praying mantises,[202] reptiles such as snakes and chameleons,[203] and birds including canaries, parakeets, and parrots[204] all finding a place. However, the most kept pet species are mammals, namely dogs, cats, and rabbits.[205][206][207] There is a tension between the role of animals as companions to humans, and their existence as individuals with rights of their own.[208] A wide variety of terrestrial and aquatic animals are hunted for sport.[209]

Symbolic uses

Artistic vision: Still Life with Lobster and Oysters by Alexander Coosemans, c. 1660
Artistic vision: Still Life with Lobster and Oysters by Alexander Coosemans, c. 1660

Animals have been the subjects of art from the earliest times, both historical, as in Ancient Egypt, and prehistoric, as in the cave paintings at Lascaux. Major animal paintings include Albrecht Dürer's 1515 The Rhinoceros, and George Stubbs's c. 1762 horse portrait Whistlejacket.[210] Insects, birds and mammals play roles in literature and film,[211] such as in giant bug movies.[212][213][214]

Animals including insects[215] and mammals[216] feature in mythology and religion. In both Japan and Europe, a butterfly was seen as the personification of a person's soul,[215][217][218] while the scarab beetle was sacred in ancient Egypt.[219] Among the mammals, cattle,[220] deer,[216] horses,[221] lions,[222] bats,[223] bears,[224] and wolves[225] are the subjects of myths and worship. The signs of the Western and Chinese zodiacs are based on animals.[226][227]

Discover more about In human culture related topics

Beef

Beef

Beef is the culinary name for meat from cattle.

Domestication of animals

Domestication of animals

The domestication of animals is the mutual relationship between non-human animals and the humans who have influence on their care and reproduction.

Livestock

Livestock

Livestock are the domesticated animals raised in an agricultural setting to provide labor and produce diversified products for consumption such as meat, eggs, milk, fur, leather, and wool. The term is sometimes used to refer solely to animals who are raised for consumption, and sometimes used to refer solely to farmed ruminants, such as cattle, sheep, goats, and pigs. Horses are considered livestock in the United States. The USDA classifies pork, veal, beef, and lamb (mutton) as livestock, and all livestock as red meat. Poultry and fish are not included in the category. The latter is likely due to the fact that fish products are not governed by the USDA, but by the FDA.

Animal husbandry

Animal husbandry

Animal husbandry is the branch of agriculture concerned with animals that are raised for meat, fibre, milk, or other products. It includes day-to-day care, selective breeding, and the raising of livestock. Husbandry has a long history, starting with the Neolithic Revolution when animals were first domesticated, from around 13,000 BC onwards, predating farming of the first crops. By the time of early civilisations such as ancient Egypt, cattle, sheep, goats, and pigs were being raised on farms.

Fishing

Fishing

Fishing is the activity of trying to catch fish. Fish are often caught as wildlife from the natural environment, but may also be caught from stocked bodies of water such as ponds, canals, park wetlands and reservoirs. Fishing techniques include hand-gathering, spearing, netting, angling, shooting and trapping, as well as more destructive and often illegal techniques such as electrocution, blasting and poisoning.

Fish farming

Fish farming

Fish farming or pisciculture involves commercial breeding of fish, most often for food, in fish tanks or artificial enclosures such as fish ponds. It is a particular type of aquaculture, which is the controlled cultivation and harvesting of aquatic animals such as fish, crustaceans, molluscs and so on, in natural or pseudo-natural environment. A facility that releases juvenile fish into the wild for recreational fishing or to supplement a species' natural numbers is generally referred to as a fish hatchery. Worldwide, the most important fish species produced in fish farming are carp, catfish, salmon and tilapia.

Cephalopod

Cephalopod

A cephalopod is any member of the molluscan class Cephalopoda such as a squid, octopus, cuttlefish, or nautilus. These exclusively marine animals are characterized by bilateral body symmetry, a prominent head, and a set of arms or tentacles modified from the primitive molluscan foot. Fishers sometimes call cephalopods "inkfish", referring to their common ability to squirt ink. The study of cephalopods is a branch of malacology known as teuthology.

Chicken

Chicken

The chicken is a domesticated junglefowl species, with attributes of wild species such as the grey and the Ceylon junglefowl that are originally from Southeast Asia. Rooster and cock are terms for adult male birds, and a younger male may be called a cockerel. A male that has been castrated is a capon. An adult female bird is called a hen, and a sexually immature female is called a pullet. Humans keep chickens primarily as a source of food or as pets. Traditionally they were also bred for cockfighting, which is still practiced in some places. Chickens domesticated for meat are broilers and for eggs are layers.

Cattle

Cattle

Cattle are large, domesticated, cloven-hooved, herbivores. They are a prominent modern member of the subfamily Bovinae and the most widespread species of the genus Bos. Adult females are referred to as cows and adult males are referred to as bulls.

Pig

Pig

The pig, often called swine, hog, or domestic pig when distinguishing from other members of the genus Sus, is an omnivorous, domesticated, even-toed, hoofed mammal. It is variously considered a subspecies of Sus scrofa or a distinct species. The pig's head-plus-body length ranges from 0.9 to 1.8 m, and adult pigs typically weigh between 50 and 350 kg, with well-fed individuals even exceeding this range. The size and weight of hogs largely depends on their breed. Compared to other artiodactyls, a pig's head is relatively long and pointed. Most even-toed ungulates are herbivorous, but pigs are omnivores, like their wild relative. Pigs grunt and make snorting sounds.

Carmine

Carmine

Carmine – also called cochineal, cochineal extract, crimson lake, or carmine lake – is a pigment of a bright-red color obtained from the aluminium complex derived from carminic acid. Specific code names for the pigment include natural red 4, C.I. 75470, or E120. Carmine is also a general term for a particularly deep-red color.

Cochineal

Cochineal

The cochineal is a scale insect in the suborder Sternorrhyncha, from which the natural dye carmine is derived. A primarily sessile parasite native to tropical and subtropical South America through North America, this insect lives on cacti in the genus Opuntia, feeding on plant moisture and nutrients. The insects are found on the pads of prickly pear cacti, collected by brushing them off the plants, and dried.

Source: "Animal", Wikipedia, Wikimedia Foundation, (2023, March 21st), https://en.wikipedia.org/wiki/Animal.

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Notes
  1. ^ Henneguya zschokkei does not have mitochondrial DNA or utilize aerobic respiration.[21]
  2. ^ The application of DNA barcoding to taxonomy further complicates this; a 2016 barcoding analysis estimated a total count of nearly 100,000 insect species for Canada alone, and extrapolated that the global insect fauna must be in excess of 10 million species, of which nearly 2 million are in a single fly family known as gall midges (Cecidomyiidae).[77]
  3. ^ Not including parasitoids.[73]
  4. ^ Compare File:Annelid redone w white background.svg for a more specific and detailed model of a particular phylum with this general body plan.
  5. ^ In his History of Animals and Parts of Animals.
  6. ^ The prefix une espèce de is pejorative.[170]
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