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Amphioctopus fangsiao

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Amphioctopus fangsiao
Octopus ocellatus (catch).jpg
Scientific classification edit
Kingdom: Animalia
Phylum: Mollusca
Class: Cephalopoda
Order: Octopoda
Family: Octopodidae
Genus: Amphioctopus
Species:
A. fangsiao
Binomial name
Amphioctopus fangsiao
(d'Orbigny, 1839)
Synonyms
  • Amphioctopus areolatus (de Haan, 1839)
  • Octopus areolatus de Haan, 1839
  • Octopus brocki Ortmann, 1888
  • Octopus fangsiao d'Orbigny, 1839
  • Octopus fangsiao etchuanus Sasaki, 1929
  • Octopus ocellatus Gray, 1849[1]

Amphioctopus fangsiao, called webfoot octopus,[2] is a species of octopus, a cephalopod belonging to the genus Amphioctopus.[3] It is found in the Pacific Ocean, including off the coasts of New Zealand[4] as well as in the Yellow Sea and surrounding Chinese shores. It is also commercially fished.[5][6]

Individuals of A. fangsiao that share genetic similarities by region are distributed throughout the Yellow Sea by size.[7][8] They can also be infected by parasites in genus Aggregata and Octopicola.[9]

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Octopus

Octopus

An octopus is a soft-bodied, eight-limbed mollusc of the order Octopoda. The order consists of some 300 species and is grouped within the class Cephalopoda with squids, cuttlefish, and nautiloids. Like other cephalopods, an octopus is bilaterally symmetric with two eyes and a beaked mouth at the center point of the eight limbs. The soft body can radically alter its shape, enabling octopuses to squeeze through small gaps. They trail their eight appendages behind them as they swim. The siphon is used both for respiration and for locomotion, by expelling a jet of water. Octopuses have a complex nervous system and excellent sight, and are among the most intelligent and behaviourally diverse of all invertebrates.

Cephalopod

Cephalopod

A cephalopod is any member of the molluscan class Cephalopoda such as a squid, octopus, cuttlefish, or nautilus. These exclusively marine animals are characterized by bilateral body symmetry, a prominent head, and a set of arms or tentacles modified from the primitive molluscan foot. Fishers sometimes call cephalopods "inkfish", referring to their common ability to squirt ink. The study of cephalopods is a branch of malacology known as teuthology.

Amphioctopus

Amphioctopus

Amphioctopus is a genus of octopuses comprising around 16 species.

Pacific Ocean

Pacific Ocean

The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean in the south, and is bounded by the continents of Asia and Oceania in the west and the Americas in the east.

New Zealand

New Zealand

New Zealand is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island and the South Island —and over 700 smaller islands. It is the sixth-largest island country by area, covering 268,021 square kilometres (103,500 sq mi). New Zealand is about 2,000 kilometres (1,200 mi) east of Australia across the Tasman Sea and 1,000 kilometres (600 mi) south of the islands of New Caledonia, Fiji, and Tonga. The country's varied topography and sharp mountain peaks, including the Southern Alps, owe much to tectonic uplift and volcanic eruptions. New Zealand's capital city is Wellington, and its most populous city is Auckland.

Yellow Sea

Yellow Sea

The Yellow Sea is a marginal sea of the Western Pacific Ocean located between mainland China and the Korean Peninsula, and can be considered the northwestern part of the East China Sea. It is one of four seas named after common colour terms, and its name is descriptive of the golden-yellow color of the silt-ridden water discharged from major rivers.

Commercial fishing

Commercial fishing

Commercial fishing is the activity of catching fish and other seafood for commercial profit, mostly from wild fisheries. It provides a large quantity of food to many countries around the world, but those who practice it as an industry must often pursue fish far into the ocean under adverse conditions. Large-scale commercial fishing is also known as industrial fishing.

Aggregata

Aggregata

Aggregata is a genus of parasitic alveolates belonging to the phylum Apicomplexa.

Distribution

This species occurs in the Philippine Sea, the northwest Pacific and off the coast of Japan (Osaka Bay).

The lifespan of this species is approximately a year, typically spawning in the spring and growing linearly after spawning.[10] In Haizhou Bay in the Yellow Sea, smaller individuals of A. fangsiao tend to be found closer to the shore in the spring and even closer during the summer, while larger individuals tend to be distributed further away from the shore.[10]

Embryonic development

A. fangsiao holds an embryonic temperature threshold in which growth accelerates at 21–24 °C (70–75 °F). In three days after gestation the octopus is typically still gastrulating. On day 5, the embryo completes their first inversion and between days 15–21 after gestation, the octopus embryo exhibit chromatophores and black eyes.[5] The embryos complete their second inversion between days 25–31.[5] Octopus embryos develop logarithmically faster with increase in temperature; those embryos who develop at a temperature of 24 °C (75 °F) go through 0.83 embryonic stages per day on average while embryos that develop at a temperature between 18 and 21 °C (64 and 70 °F) go through 0.67 embryonic stages per day on average.[5] The octopuses that had a smaller embryo but larger yolk volume tended to hatch at 24 °C (75 °F).[5]

Abnormal morphologies

The embryos that grew at 27 °C (81 °F) were not able to hatch due to accelerated growth which caused eggs to be more swollen than eggs developing at lower temperatures; this accelerated growth caused the chorion to break and the embryo to hatch prematurely.[5] While the eggs grown at 21 °C (70 °F) were ovular in shape as normal, the eggs grown at 24 °C (75 °F) were malformed.[5] The embryos grown in 24 °C (75 °F) conditions also exhibited red eyes at 2 days after gestation.[5]

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Gestation

Gestation

Gestation is the period of development during the carrying of an embryo, and later fetus, inside viviparous animals. It is typical for mammals, but also occurs for some non-mammals. Mammals during pregnancy can have one or more gestations at the same time, for example in a multiple birth.

Gastrulation

Gastrulation

Gastrulation is the stage in the early embryonic development of most animals, during which the blastula, or in mammals the blastocyst is reorganized into a multilayered structure known as the gastrula. Before gastrulation, the embryo is a continuous epithelial sheet of cells; by the end of gastrulation, the embryo has begun differentiation to establish distinct cell lineages, set up the basic axes of the body, and internalized one or more cell types including the prospective gut.

Embryo

Embryo

An embryo is an initial stage of development of a multicellular organism. In organisms that reproduce sexually, embryonic development is the part of the life cycle that begins just after fertilization of the female egg cell by the male sperm cell. The resulting fusion of these two cells produces a single-celled zygote that undergoes many cell divisions that produce cells known as blastomeres. The blastomeres are arranged as a solid ball that when reaching a certain size, called a morula, takes in fluid to create a cavity called a blastocoel. The structure is then termed a blastula, or a blastocyst in mammals.

Inversion (evolutionary biology)

Inversion (evolutionary biology)

In evolutionary developmental biology, inversion refers to the hypothesis that during the course of animal evolution, the structures along the dorsoventral (DV) axis have taken on an orientation opposite that of the ancestral form.

Chromatophore

Chromatophore

Chromatophores are cells that produce color, of which many types are pigment-containing cells, or groups of cells, found in a wide range of animals including amphibians, fish, reptiles, crustaceans and cephalopods. Mammals and birds, in contrast, have a class of cells called melanocytes for coloration.

Yolk

Yolk

Among animals which produce eggs, the yolk is the nutrient-bearing portion of the egg whose primary function is to supply food for the development of the embryo. Some types of egg contain no yolk, for example because they are laid in situations where the food supply is sufficient or because the embryo develops in the parent's body, which supplies the food, usually through a placenta. Reproductive systems in which the mother's body supplies the embryo directly are said to be matrotrophic; those in which the embryo is supplied by yolk are said to be lecithotrophic. In many species, such as all birds, and most reptiles and insects, the yolk takes the form of a special storage organ constructed in the reproductive tract of the mother. In many other animals, especially very small species such as some fish and invertebrates, the yolk material is not in a special organ, but inside the egg cell.

Diseases affecting A. fangsiao

Like many other species of cephalopods, A. fangsiao is not safe from infection. As of 2021, there is not much information on the protozoan parasites that infect this species among other species of cephalopods native to the Pacific Ocean. It is known that is that these parasites belong to the genus Aggregata and are responsible for the infections that have led to the death of many A. fangsiao in a culture facility in China.[11] Infection in A. fangsiao was also found from the genus Octopicola.[11]

Aggregata infection

Evidence of the parasite's infection on the A. fangsiao specimens include numerous white cysts scattered throughout the body of each octopus.[11] These cysts, called sporocysts, are smooth to the touch, measuring 17.69–20.72 μm by 15.97–20.00 μm. While the oocysts themselves are circular in shape measuring 249.75–501.75 μm and 116.84–350.87 μm in size.[11] Oocysts represent one of the stages in their lifecycle.[11] When histological surveys were done, the cysts had severely affected the integrity of the internal organs, and often the intestinal issue was greatly affected by oocysts filled with sporocysts.[11] Out of the 220 octopuses collected at the site, 95 of them were infected with Aggregata—representing 43% of the investigated specimens in the facility.[11]

Octopicola huanghaiensis infection

Recently discovered, the species Octopicola huanghaiensis is the first of its genus to infect A. fangsiao.[12] However, its genus, Octopicola, is known to commonly infect octopuses.[12]

Abnormalities within infection rates

Another experiment looked at the rate of infection of females versus males.[11] The results were not as expected, 64% of the fatalities were female while the remaining 36% were males.[11] The sample of the collected deceased A. fangsiao was 1,303; these data were collected between March 16 to April 2, 2017, in the Yellow Sea.[11] These data showed that the death rate of females was higher than males during their breeding season.[11] Upon further examination of the dead female specimens the researchers found that 16% had edema, 37% had broken skin, and 4% had broken arms.[11]

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Protozoan infection

Protozoan infection

Protozoan infections are parasitic diseases caused by organisms formerly classified in the kingdom Protozoa. They are usually contracted by either an insect vector or by contact with an infected substance or surface and include organisms that are now classified in the supergroups Excavata, Amoebozoa, SAR, and Archaeplastida.

Aggregata

Aggregata

Aggregata is a genus of parasitic alveolates belonging to the phylum Apicomplexa.

Cyst

Cyst

A cyst is a closed sac, having a distinct envelope and division compared with the nearby tissue. Hence, it is a cluster of cells that have grouped together to form a sac ; however, the distinguishing aspect of a cyst is that the cells forming the "shell" of such a sac are distinctly abnormal when compared with all surrounding cells for that given location. A cyst may contain air, fluids, or semi-solid material. A collection of pus is called an abscess, not a cyst. Once formed, a cyst may resolve on its own. When a cyst fails to resolve, it may need to be removed surgically, but that would depend upon its type and location.

Histology

Histology

Histology, also known as microscopic anatomy or microanatomy, is the branch of biology that studies the microscopic anatomy of biological tissues. Histology is the microscopic counterpart to gross anatomy, which looks at larger structures visible without a microscope. Although one may divide microscopic anatomy into organology, the study of organs, histology, the study of tissues, and cytology, the study of cells, modern usage places all of these topics under the field of histology. In medicine, histopathology is the branch of histology that includes the microscopic identification and study of diseased tissue. In the field of paleontology, the term paleohistology refers to the histology of fossil organisms.

Seasonal breeder

Seasonal breeder

Seasonal breeders are animal species that successfully mate only during certain times of the year. These times of year allow for the optimization of survival of young due to factors such as ambient temperature, food and water availability, and changes in the predation behaviors of other species. Related sexual interest and behaviors are expressed and accepted only during this period. Female seasonal breeders will have one or more estrus cycles only when she is "in season" or fertile and receptive to mating. At other times of the year, they will be anestrus, or have a dearth of their sexual cycle. Unlike reproductive cyclicity, seasonality is described in both males and females. Male seasonal breeders may exhibit changes in testosterone levels, testes weight, and fertility depending on the time of year.

Edema

Edema

Edema, also spelled oedema, and also known as fluid retention, dropsy, hydropsy and swelling, is the build-up of fluid in the body's tissue. Most commonly, the legs or arms are affected. Symptoms may include skin which feels tight, the area may feel heavy, and joint stiffness. Other symptoms depend on the underlying cause.

Genetic structure

These octopuses are generally found along the shores of China, and are relatively highly genetic diverse in haplotype diversity, especially in the Zhanjiang province while the lowest haplotype diversity is seen in the Qingdao province. They also have low nucleotide diversity.[7] There is more genetic variation among the population of A. fansgsiao than there are within the population of A. fangsiao.[7]

Two groups were defined by their genetic similarities and by region: A. fangsiao in the northern shore, including Dalian, Yantai, Qingdao, and Nantong provinces, were more similar among each other while the A. fangsiao in the southern shore, including Wenzhou, Xiamen, Dongshan, and Huizhou provinces, were more genetically similar each other.[7] A haplotype network analysis showed that all the octopuses in the northern provinces shared a haplotype with each other using several mitochondrial gene markers (ATPase 6, ND2, and ND5), while the octopuses in the southern provinces shared different haplotypes with each gene marker; in all gene markers, the northern and southern province octopuses did not share any haplotypes with each other.[7]

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China

China

China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and borders fourteen countries by land, the most of any country in the world, tied with Russia. With an area of approximately 9.6 million square kilometres (3,700,000 sq mi), it is the world's third largest country by total land area. The country consists of 22 provinces, five autonomous regions, four municipalities, and two special administrative regions. The national capital is Beijing, and the most populous city and largest financial center is Shanghai.

Haplotype

Haplotype

A haplotype is a group of alleles in an organism that are inherited together from a single parent.

Qingdao

Qingdao

Qingdao is a major city in eastern Shandong Province. The city's name in Chinese characters literally means "azure island". Located on China's Yellow Sea coast, it is a major nodal city of the Belt and Road Initiative (BRI) that connects Asia with Europe. It has the highest GDP of any city in the province. Administered at the sub-provincial level, Qingdao has jurisdiction over seven districts and three county-level cities. As of the 2020 census, Qingdao built-up area made of the 7 urban Districts was home to 7,172,451 inhabitants making it the 15th largest city in China by population. Lying across the Shandong Peninsula and looking out to the Yellow Sea, it borders the prefecture-level cities of Yantai to the northeast, Weifang to the west and Rizhao to the southwest.

Nucleotide diversity

Nucleotide diversity

Nucleotide diversity is a concept in molecular genetics which is used to measure the degree of polymorphism within a population.

Genetic variation

Genetic variation

Genetic variation is the difference in DNA among individuals or the differences between populations. The multiple sources of genetic variation include mutation and genetic recombination. Mutations are the ultimate sources of genetic variation, but other mechanisms, such as genetic drift, contribute to it, as well.

Dalian

Dalian

Dalian is a major sub-provincial port city in Liaoning province, People's Republic of China, and is Liaoning's second largest city and the third-most populous city of Northeast China. Located on the southern tip of Liaodong peninsula, it is the southernmost city in both Liaoning and the entire Northeast. Dalian borders the prefectural cities of Yingkou and Anshan to the north and Dandong to the northeast, and also shares maritime boundaries with Qinhuangdao and Huludao across the Liaodong Bay to west and northwest, Yantai and Weihai on the Shandong peninsula across the Bohai Strait to the south, and North Korea across the Korea Bay to the east.

Nantong

Nantong

Nantong is a prefecture-level city in southeastern Jiangsu province, China. Located on the northern bank of the Yangtze River, near the river mouth. Nantong is a vital river port bordering Yancheng to the north; Taizhou to the west; Suzhou, Wuxi and Shanghai to the south across the river; and the East China Sea to the east. Its population was 7,726,635 as of the 2020 census, 3,766,534 of whom lived in the built-up area made up of three urban districts.

Wenzhou

Wenzhou

Wenzhou, historically known as Wenchow is a prefecture-level city in southeastern Zhejiang province in the People's Republic of China. Wenzhou is located at the extreme south east of Zhejiang Province with its borders connecting to Lishui on the west, Taizhou on the north, and Fujian to the south. It is surrounded by mountains, the East China Sea, and 436 islands, while its lowlands are almost entirely along its East China Sea coast, which is nearly 355 kilometres in length. Most of Wenzhou's area is mountainous as almost 76 percent of its 11,784-square-kilometre (4,550 sq mi) surface area is classified as mountains and hills. It is said that Wenzhou has 7/10 mountains, 1/10 water, and 2/10 farmland. At the time of the 2010 Chinese census, 3,039,500 people lived in Wenzhou's urban area; the area under its jurisdiction held a population of 9,122,100 of which 31.16% are non-local residents from outside of Wenzhou.

Xiamen

Xiamen

Xiamen, also known as Amoy, is a sub-provincial city in southeastern Fujian, People's Republic of China, beside the Taiwan Strait. It is divided into six districts: Huli, Siming, Jimei, Tong'an, Haicang, and Xiang'an. All together, these cover an area of 1,700.61 square kilometers (656.61 sq mi) with a population of 5,163,970 as of 2020 and estimated at 5.308 million as of 31 December 2022. The urbanized area of the city has spread from its original island to include most parts of all six of its districts, and with 4 Zhangzhou districts, form a built-up area of 7,284,148 inhabitants. This area also connects with Quanzhou in the north, making up a metropolis of nearly ten million people. The Kinmen Islands (Quemoy) administered by the Republic of China (Taiwan) which lie less than 6 kilometers (4 mi) away separated by Xiamen Bay. As part of the Opening Up Policy under Deng Xiaoping, Xiamen became one of China's original four special economic zones opened to foreign investment and trade in the early 1980s.

Dongshan County

Dongshan County

Dongshan (help·info) is a county of far southern Fujian Province, People's Republic of China, located along the Taiwan Strait. It comprises 44 islands for a total area of 194 square kilometres (75 sq mi) and is under the administration of Zhangzhou City. The total population was 200,000. Dongshan County has jurisdiction over seven towns, a nationally managed forest and an economic and technological development district. It is an important port for international trade and trade with Taiwan.

Huizhou

Huizhou

Huizhou is a city in central-east Guangdong Province, China, forty-three miles north of Hong Kong. Huizhou borders the provincial capital of Guangzhou to the west, Shenzhen and Dongguan to the southwest, Shaoguan to the north, Heyuan to the northeast, Shanwei to the east, and Daya Bay of the South China Sea to the south. As of the 2020 census, the city has about 6,042,852 inhabitants and is administered as a prefecture-level city. Huizhou's core metropolitan area, which is within Huicheng and Huiyang Districts, is home to around 2,090,578 inhabitants.

Mitochondrial DNA

Mitochondrial DNA

Mitochondrial DNA is the DNA located in mitochondria, cellular organelles within eukaryotic cells that convert chemical energy from food into a form that cells can use, such as adenosine triphosphate (ATP). Mitochondrial DNA is only a small portion of the DNA in a eukaryotic cell; most of the DNA can be found in the cell nucleus and, in plants and algae, also in plastids such as chloroplasts.

Culinary use

In Korea, Amphioctopus fangsiao is called jukkumi (주꾸미) or jjukkumi (쭈꾸미) and is often stir-fried in spicy gochujang-based sauce.[13][14]

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Korea

Korea

Korea is a peninsular region in East Asia. Since 1945, it has been divided at or near the 38th parallel, with North Korea comprising its northern half and South Korea comprising its southern half. Korea consists of the Korean Peninsula, Jeju Island, and several minor islands near the peninsula. The peninsula is bordered by China (Manchuria) to the north and Russia to the northeast, across the Amrok and Duman rivers. It is separated from Japan to the southeast by the Korea Strait.

Gochujang

Gochujang

Gochujang or red chili paste is a savory, sweet, and spicy fermented condiment popular in Korean cooking. It is made from gochu-garu, glutinous rice, meju powder, yeotgireum, and salt. The sweetness comes from the starch of cooked glutinous rice, cultured with saccharifying enzymes during the fermentation process. Traditionally, it has been naturally fermented over years in jangdok (earthenware) on an elevated stone platform, called jangdokdae, in the backyard.

Bokkeum

Bokkeum

Bokkeum (볶음) is a category of stir-fried dishes in Korean cuisine.

Gui (food)

Gui (food)

In Korean cuisine gui or guee is a grilled dish. Gui most commonly has meat or fish as the primary ingredient, but may in some cases also have grilled vegetables or other vegetarian ingredients. The term derives from the verb gupda, which literally means "grill". At traditional restaurants, meats are cooked at the center of the table over a charcoal grill, surrounded by various banchan and individual rice bowls. The cooked meat is then cut into small pieces and wrapped with fresh lettuce leaves, with rice, thinly sliced garlic, ssamjang, and other seasonings. The suffix gui is often omitted in the names of meat-based gui such as galbi, whose name was originally galbi gui.

Source: "Amphioctopus fangsiao", Wikipedia, Wikimedia Foundation, (2023, February 15th), https://en.wikipedia.org/wiki/Amphioctopus_fangsiao.

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References
  1. ^ "UNESCO-IOC Register of Marine Organisms (URMO) - Amphioctopus fangsiao (d'Orbigny, 1839)". Marinespecies.org. 21 October 2010. Retrieved 27 August 2011.
  2. ^ "Webfoot octopus". FishSource. 22 May 2013. Retrieved 22 May 2017.
  3. ^ "WoRMS - World Register of Marine Species - Amphioctopus fangsiao (d'Orbigny, 1839)". Marinespecies.org. 21 October 2010. Retrieved 27 August 2011.
  4. ^ "Global Barcode Of Life Data Mirror". Nz.boldmirror.net. Archived from the original on 24 February 2014. Retrieved 27 August 2011.
  5. ^ a b c d e f g h Jiang, Dianhang; Zheng, Xiaodong; Qian, Yaosen; Zhang, Qingqi (May 2020). "Embryonic development of Amphioctopus fangsiao under elevated temperatures: Implications for resource management and conservation". Fisheries Research. 225: 105479. doi:10.1016/j.fishres.2019.105479. S2CID 214538232.
  6. ^ Pang, Yumeng; Tian, Yongjun; Fu, Caihong; Ren, Yiping; Wan, Rong (1 October 2020). "Growth and Distribution of Amphioctopus fangsiao (d'Orbigny, 1839–1841) in Haizhou Bay, Yellow Sea". Journal of Ocean University of China. 19 (5): 1125–1132. Bibcode:2020JOUC...19.1125P. doi:10.1007/s11802-020-4322-7. ISSN 1993-5021. S2CID 221401494.
  7. ^ a b c d e Muhammad, Faiz; Chen, Wei; Liu, Liqin; Gong, Li; Du, Xun; Shafi, Muhammad; Lü, Zhen-ming (1 July 2019). "Genetic structure of Amphioctopus fangsiao (Mollusca, Cephalopoda) in Chinese waters inferred from variation in three mtDNA genes (ATPase 6, ND2, and ND5)". Hydrobiologia. 838 (1): 111–119. doi:10.1007/s10750-019-03981-9. ISSN 1573-5117. S2CID 254541630.
  8. ^ Pang, Yumeng; Tian, Yongjun; Fu, Caihong; Ren, Yiping; Wan, Rong (1 October 2020). "Growth and Distribution of Amphioctopus fangsiao (d'Orbigny, 1839–1841) in Haizhou Bay, Yellow Sea". Journal of Ocean University of China. 19 (5): 1125–1132. Bibcode:2020JOUC...19.1125P. doi:10.1007/s11802-020-4322-7. ISSN 1993-5021. S2CID 221401494.
  9. ^ Ren, Jing; Zheng, Xiaodong; Qian, Yaosen; Zhang, Qingqi (2019), Gestal, Camino; Pascual, Santiago; Guerra, Ángel; Fiorito, Graziano (eds.), "Pathogens and Related Diseases in Non-European Cephalopods: Asia. A Preliminary Review", Handbook of Pathogens and Diseases in Cephalopods, Cham: Springer International Publishing, pp. 227–230, doi:10.1007/978-3-030-11330-8_18, ISBN 978-3-030-11329-2
  10. ^ a b Pang, Yumeng; Tian, Yongjun; Fu, Caihong; Ren, Yiping; Wan, Rong (1 October 2020). "Growth and Distribution of Amphioctopus fangsiao (d'Orbigny, 1839–1841) in Haizhou Bay, Yellow Sea". Journal of Ocean University of China. 19 (5): 1125–1132. Bibcode:2020JOUC...19.1125P. doi:10.1007/s11802-020-4322-7. ISSN 1993-5021. S2CID 221401494.
  11. ^ a b c d e f g h i j k l Ren, Jing; Zheng, Xiaodong; Qian, Yaosen; Zhang, Qingqi (2019), Gestal, Camino; Pascual, Santiago; Guerra, Ángel; Fiorito, Graziano (eds.), "Pathogens and Related Diseases in Non-European Cephalopods: Asia. A Preliminary Review", Handbook of Pathogens and Diseases in Cephalopods, Cham: Springer International Publishing, pp. 227–230, doi:10.1007/978-3-030-11330-8_18, ISBN 978-3-030-11329-2
  12. ^ a b Du, Xin; Dong, Chao; Sun, Shi-Chun (November 2018). "Octopicola huanghaiensis n. sp. (Copepoda: Cyclopoida: Octopicolidae), a new parasitic copepod of the octopuses Amphioctopus fangsiao (d'Orbigny) and Octopus minor (Sasaki) (Octopoda: Octopodidae) in the Yellow Sea". Systematic Parasitology. 95 (8–9): 905–912. doi:10.1007/s11230-018-9819-8. ISSN 0165-5752. PMID 30276653. S2CID 255067189.
  13. ^ Chakraborty, Shruti (3 March 2016). "Seoul Food: Hitting the streets in search of Octopus". The Indian Express. Retrieved 13 April 2017.
  14. ^ Lee, Cecilia Hae-Jin (30 April 2015). "All-you-can-eat Korean BBQ at Jjukku Jjukku". Los Angeles Times. Retrieved 13 April 2017.
Further reading
  • Norman M.D. & Hochberg F.G. (2005) The current state of Octopus taxonomy. Phuket Marine Biological Center Research Bulletin 66:127–154.
  • Furuya, H. (2006). Three new species of dicyemid mesozoans (Phylum Dicyemida) from Amphioctopus fangsiao (Mollusca: Cephalopoda), with comments on the occurrence patterns of dicyemids. Zoological Science 23: 105–119.
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